BULLETIN

ISLAND LISTS OF WEST INDIAN AMPHIBIANS AND REPTILES

Robert Powell and Robert W. Henderson, Editors

Vol. 51, No. 2, pp. 85–166 2012

UNIVERSITY OF FLORIDA GAINESVILLE

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 ISLAND LISTS OF WEST INDIAN
AMPHIBIANS AND REPTILES

Robert Powell1 and Robert W. Henderson2, Editors

TABLE OF CONTENTS

Introduction by Robert Powell and Robert W. Henderson......................................86

Swan Islands by Robert Powell and Robert W. Henderson.....................................91
Bahama Islands and Turks & Caicos Islands by Sandra D. Buckner,
Richard Franz, and R. Graham Reynolds...........................................................93
Cayman Islands by Arthur C. Echternacht............................................................. 111
The Cuban Archipelago by Alberto R. Estrada...................................................... 113
Jamaica by Byron S. Wilson, Thera Edwards, and Robert Powell........................126
Hispaniola and Navassa by Robert Powell............................................................129
Puerto Rico and the Virgin Islands by Gregory C. Mayer.....................................136
Lesser Antilles by Robert W. Henderson and Michel Breuil.................................148
Literature Cited......................................................................................................160

1
Department of Biology, Avila University, Kansas City, Missouri 64145-1698, USA <[email protected]>
2
Section of Vertebrate Zoology, Milwaukee Public Museum, Milwaukee, Wisconsin 53233, USA <[email protected]>

Powell, R., and R. W. Henderson, Eds. 2012. Island lists of West Indian amphibians and reptiles. Bulletin of the Florida Museum of Natural History 51(2):85–
166.
86 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

INTRODUCTION

Robert Powell1 and Robert W. Henderson2

We believe that monitoring and documenting (1977), based largely on the Schwartz and Thomas
changes in the composition and distribution of checklist, published a paper wherein they listed
the West Indian herpetofauna is necessary and the species of frogs, turtles, lizards, snakes, and
important. This vast complex of islands with crocodilians for each of the 627 islands then known
disparate geological origins and biogeographic to harbor amphibians and reptiles. Subsequently,
relationships with one another and the mainland Schwartz and Henderson (1988, 1991) provided an
Americas supports exceptionally abundant, diverse, updated checklist and an overview of West Indian
and largely endemic populations of terrestrial herpetology, and Powell et al. (1996) published
amphibians and reptiles, and is ideally suited for an abbreviated list, each documenting substantive
studies that provide new insights into evolutionary changes in our understanding of the diversity and
and ecological relationships. In addition, we are distribution of the region’s herpetofauna. Most
motivated by increasingly critical conservation recently, Powell and Henderson (1999, 2003)
concerns in the region, which was identified as published two addenda to Powell et al. (1996)
a biodiversity hotspot by Myers et al. (2000). A and S. Blair Hedges (2012) provided a regularly
large majority of the islands are changing rapidly, updated list of species with distribution maps
resulting in dramatic alterations of habitats, nearly (http://caribherp.org) — but no updated island-by-
all of them mediated by human activities. island list of West Indian amphibians and reptiles
Thomas Barbour (1914, 1930, 1935, 1937) has appeared since the publication of MacLean et
effectively initiated the documentation of the al. 35 years ago.
region’s herpetofaunas by providing a series of Although many of the documented changes
increasingly comprehensive checklists. Although reflect a better and more modern understanding of
replete with inconsistencies reflecting the limited relationships among the islands’ species and the
and localized research of those days, now outdated resultant adjustments in taxonomy, many questions
taxonomy, and data often based on anecdotal remain. For example, at least some widely
accounts, very short visits, and hastily formed distributed species almost certainly represent
impressions (Henderson & Powell 2005), they species complexes, with populations in different
collectively constitute a baseline for comparisons habitats on large islands or on different islands or
with more modern efforts. island groups worthy of recognition as full species.
Thirty-seven years ago, Albert Schwartz and For example, currently recognized Hispaniolan
Richard Thomas (1975) provided the first scholarly and Bahamian subspecies of Anolis distichus
effort to document amphibian and reptilian diversity might well be distinct at the species level (e.g., Ng
in the West Indies when they published what has & Glor, 2011; Glor & Laport, in press), and Lesser
been hailed (Pregill & Crother 1999) as “the most Antillean populations of Thecadactylus probably
valuable contribution to the biogeography of West represent several species-level taxa (Köhler &
Indian amphibians and reptiles in the past 50 years.” Vesely, 2011).
This detailed, carefully researched checklist of the However, most documented changes illustrate
herpetofauna listed each species and the island(s) on the impact of human activities (e.g., Henderson
which it occurred. Two years later, MacLean et al. & Powell 2001, 2009). The herpetofaunas of the

1
Department of Biology, Avila University, Kansas City, Missouri 64145-1698, USA <[email protected]>
2
Section of Vertebrate Zoology, Milwaukee Public Museum, Milwaukee, Wisconsin 53233, USA <[email protected]>
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 87

West Indies are in flux; human population growth environment and, ultimately, to the resources
takes its toll on the environment as land that once necessary to sustain many species of frogs and
sustained forests is cleared for homes, agriculture, reptiles. In addition to the reduction or elimination
and charcoal production. Humans bring with them of critical resources, development for the tourist
cats and dogs that prey on reptiles, and goats, industry often entails bringing supplies (e.g.,
burros, and cattle that crop vegetation to bare soil, lumber, decorative plants) largely from the United
eliminating critical ground cover and food for a States, mainland South America, or other islands,
variety of amphibians and reptiles. The mongoose and non-native herpetofaunal species sometimes
was introduced to many islands in the 19th century, arrive with them as stowaways (e.g., Kraus 2009;
putatively to control rodent populations that were Powell et al. 2011). For example, in recent years,
having a deleterious effect on sugar production. Anolis carolinensis (native to the U.S.) has arrived
The impact on ground-dwelling lizards, snakes, on Anguilla, A. sagrei (native to Cuba and the
and birds has frequently been catastrophic. Bahamas but very common in most of Florida) has
With many island economies increasingly become established on Grenada, St. Vincent, and
based on tourism, development is rampant; with St. Maarten, and A. cristatellus (native to the Puerto
development come radical modifications to the Rico Bank) has invaded the Dominican Republic

Figure 1. Major islands or islands groups of the West Indies used in this study.
88 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

and Dominica, and has been reported from St. although this sometimes places geographically
Maarten. Similarly, the large Cuban treefrog, proximate islands far apart in the lists. We provide
Osteopilus septentrionalis, has become established latitude and longitude for most islands (although
in the Virgin Islands and Lesser Antilles, and rounding decimal degrees to the nearest hundredth
the snake, Pantherophis guttatus (native to the will provide only an approximate location for
eastern U.S.), has been captured in the Bahamian many small cays). We also include area for most
Archipelago, the U.S. Virgin Islands, and several islands, and maps that illustrate the geographic
islands in the Lesser Antilles. relationships of major islands. Except for Cayo
In collaboration with experts most familiar Arenas (Cayos Siete Hermanos, Hispaniola, which
with specific areas within the West Indies, we set we have surveyed), we exclude islands for which
out to generate new island-by-island lists. Our no records exist, although we cannot, in most
reasons for doing so are fivefold: Since 1977, instances, distinguish between islands that have
(1) we have found the (now outdated) island list been surveyed without finding any resident species
by MacLean et al. to be a useful tool; (2) a great (e.g., Cayo Arenas) and those that have not been
deal of new information on the distribution of surveyed.
amphibians and reptiles on West Indian islands has Records are based on museum vouchers and
accumulated; (3) many new species of West Indian localities cited in the literature supplemented by the
frogs and reptiles have been described; (4) several authors’ observations and a few sightings by reliable
extirpations (or possible extinctions) have occurred; witnesses. We generally follow the taxonomy
and (5) a number of species (mostly from the U.S.) provided on CaribHerp (Hedges 2012). Two notable
have been introduced to West Indian islands and exceptions are the inclusion of currently recognized
might ultimately have a deleterious impact on the subspecies (in order to enhance the utility of this
native herpetofauna. list in case some of these are elevated to species in
the future) and the choice not to elevate Bahamian
THE LISTS
subspecies of Cyclura to full species. Although we
We have defined the West Indies to include the Swan are inclined to believe that elevation is appropriate,
Islands, Cayman Islands, Bahama Islands, Greater that decision must be based on detailed genetic data
Antilles (Cuba, Jamaica, Hispaniola, and Greater that are not yet available; consequently, we take
Puerto Rico), and Lesser Antilles (extending from a conservative approach. We denote species that
Sombrero in the north to Grenada in the south). The have been introduced to an island with an “(I).” We
following lists document more than 700 species do not distinguish between newly introduced taxa
of amphibians and reptiles that occur on over 700 and presumed reintroductions, mainly because the
islands. They are organized first by major island historical presence of some populations cannot
(e.g., Cuba) or island group (e.g., Lesser Antilles), be confirmed. Furthermore, the origins of some
then by island bank (e.g., Grenada Bank in the species cannot be established. Consequently,
Lesser Antilles). These are listed in a roughly west- our annotations frequently are accompanied by
to-east fashion for the Greater Antilles and north- question marks. For example, populations of
to-south for the Lesser Antilles. This geographic Chelonoidis carbonaria, Iguana iguana, and
approach is largely retained for major islands or human commensals like Hemidactylus mabouia on
island groups within an island bank (defined for many, especially Lesser Antillean islands, likely are
our purposes as a contiguous emergent land mass comprised of descendants of animals that arrived
during the Pleistocene glacial maxima regardless by natural means, descendants of individuals
of how many individual islands are extant today). transported to the islands by the Amerindians who
Satellites to each of the major islands or within first colonized these islands some 7,000 years ago,
island groups are listed alphabetically to facilitate and animals recently transported to the islands,
finding them in what are frequently long lists, either inadvertently accompanying shipments
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 89

of goods or intentionally as pets (e.g., Iverson some species may have become extinct, some
1978; Censky 1988; Greene et al. 2002; Powell populations certainly have been extirpated, and
et al. 2011). For I. iguana, we believe that we can many undoubtedly are threatened or endangered
with some accuracy distinguish natural (endemic) (e.g., Hedges 1999; Stuart et al. 2004; Henderson
populations from those that are introduced, and we & Powell 2009; IUCN 2011) — but assessments
have marked them accordingly. For C. carbonaria on our part at this time would have to have been
and H. mabouia, we consistently use “(I?)” (except based almost entirely on circumstantial evidence,
for two populations of the former that are known since detailed studies for most species are lacking.
to be introduced by human agency in the British Occasionally, records have been published
Virgin Islands and one of the latter that was recently about specimens or fossil remains (e.g., tortoises
introduced in the Turks and Caicos Islands), although in the genus Chelonoidis) for which the species
we are sure that some populations, especially in identity is unknown or uncertain. We have listed
the southern Lesser Antilles, arrived naturally and these accordingly (i.e., Chelonoidis sp.). Also,
many, especially in the Greater Antilles, are almost in a few instances, the assignment of a particular
certainly introduced. Also, Rhinella marina was island population to species or (more commonly)
intentionally introduced onto many West Indian subspecies is uncertain (e.g., Celestus stenurus ssp.
islands (e.g., Lever 2003; Powell et al. 2011), but on Île Grande Cayemite off Hispaniola) or an island
populations on some of the southern Lesser Antilles population that is likely to represent a new species
might have arrived by natural over-water dispersal yet to be formally described (e.g., Sphaerodactylus
(Henderson & Powell 2009). We mark the latter sp. on Sombrero).
with “(I?).” Complicating matters even further are Until recently, the systematics of West Indian
species that have been documented from islands skinks has been problematic. Except for Mabuya
but are not known to have established breeding lineolatus from Hispaniola and M. macleani from
populations. We have designated these as waifs Carrot Rock, British Virgin Islands, relationships
and annotated such listings with a “(W)”; when we were poorly understood (e.g., Mayer & Lazell
have data that suggest a population designated as 2000). For most of the last century, all other island
a waif might be established, we mark these with a populations from the region had been assigned to M.
“(W?).” mabouya (e.g., Dunn 1936; Schwartz & Henderson
Although documenting a negative is 1991). More recently, Henderson & Powell (2009)
impossible, we have endeavored to identify designated those populations as “Mabuya sp.”
populations that are believed to have been Herein we adopt the taxonomy of Hedges and
extirpated by marking them with a “(PX)” (= Conn (2012), who recognized six genera (Alinea,
possibly or probably extirpated). Similarly, we Capitellum, Copeoglossum, Mabuya, Marisora,
have marked presumably extinct taxa with a and Spondylurus) and 33 species in the West Indies,
“(PE)” (= possibly extinct). Taxa known from an 16 of which are possibly extinct.
island only from the fossil record are indicated Certainly, like the ever-changing taxonomy,
with an “(F),” whereas extant taxa for which fossils all of the data presented herein are inevitably
are known are indicated with an “(F*).” We also subject to change as human populations continue
considered attempts to document the conservation to grow, habitats are increasingly altered, and new
status of species (e.g., threatened or endangered), introductions of alien species occur — but also as
but ultimately chose not to do so, mainly because new species are described and new populations
the status of many populations is uncertain and we are discovered. Much of the attraction for
did not want to imply that circumstances were less herpetologists working in the West Indies is the
than critical by omitting annotations of status. For vast array of unanswered questions, the answers
example, the status of most West Indian reptiles has to many of which will undoubtedly elude us for
not been assessed (e.g., Henderson & Powell 2009); generations to come.
90 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

ACKNOWLEDGMENTS and the reviewers for helpful comments. Lourdes

Rodríguez Schettino, editor of seminal works on
The Falconwood Foundation, through a grant the Cuban herpetofauna (1999; 2003), went well
to The Conservation Agency, provided financial beyond the call of duty for reviewers by providing
support for this project. Recent West Indian field island sizes and current information, much of
work of RWH has been generously funded by the it unpublished, on the distribution of Cuban
Windway Foundation and the Milwaukee Public amphibians and reptiles.
Museum, and that of RP by a series of National Rose Henderson (1945–2012) accompanied
Science Foundation “Research Experiences for her husband RWH during his first trips to the
Undergraduates” grants and Avila University. Lesser Antilles. We dedicate this publication to her
In addition to the contributors to this effort, memory.
we thank John S. Parmerlee, Jr. for providing maps

Rose Henderson during an ascent of La Soufrière

on Guadeloupe in 1987.
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 91

SWAN ISLANDS

Robert Powell1 and Robert W. Henderson2

The Swan Islands consist of three “major” islands a female (MCZ 11979) donated to the museum in
(including Booby Cay [= El Cayo Pájaro Bobo], 1916. We have deleted Cnemidophorus lemniscatus
with an area of <0.01 km2) and several even from our list (following Schwartz & Henderson
smaller satellites. Herpetological records exist 1988 and subsequent checklists), but tentatively
only for Great Swan and Little Swan islands. Nine retain A. ameiva fuliginosa and Cubophis brooksi.
species of reptiles have been documented, but the (formerly considered a subspecies of Cubophis
lack of recent surveys and the questionable nature cantherigerus.)
of several records suggest that only six species are
present. Ameiva ameiva fuliginosa has not been ACKNOWLEDGMENTS
collected on Great Swan Island since 1884; Barbour
(1914) reported that this population was extirpated Larry David Wilson, James R. McCranie, and
and Schwartz and Henderson (1988) suggested Josiah Townsend provided valuable insights on an
that the original record might have been a locality early draft of this list.
error. Schwartz and Thomas (1975) and MacLean
et al. (1977) listed Cnemidophorus lemniscatus SWAN ISLAND BANK (= Islas del Cisne/Islas
lemniscatus from Great Swan Island, but we can Santanilla)
Great Swan Island (= Cisne Grande/Isla Grande)
find no other records of the species from the islands.
(17.41°/-83.93°) (5.5 km²)
Cubophis brooksi was described on the basis of Ameiva ameiva fuliginosa (PX)
two specimens (MCZ 7893) by Barbour (1914), Anolis nelsoni
and Kardong (1980) described the musculature of Aristelliger nelsoni

Figure 2. Map of the Swan Islands.

1
Department of Biology, Avila University, Kansas City, Missouri 64145-1698, USA <[email protected]>
2
Section of Vertebrate Zoology, Milwaukee Public Museum, Milwaukee, Wisconsin 53233, USA <[email protected]>
92 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Iguana iguana
Leiocephalus varius
Sphaerodactylus exsul
Epictia magnamaculata
Little Swan Island (= Cisne Pequeño/Isla Pequeño)
(17.41°/-83.90°) (2.5 km²)
Anolis nelsoni
Aristelliger nelsoni
Iguana iguana
Leiocephalus varius
Sphaerodactylus exsul
Cubophis brooksi
Epictia magnamaculata
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 93

BAHAMA ISLANDS AND TURKS & CAICOS ISLANDS

Sandra D. Buckner1, Richard Franz2, and R. Graham Reynolds3

The Bahamian Archipelago lies in the North Atlan- The Bahamian Archipelago is divided into
tic, southeast of Florida and north of Cuba and two political identities. The Commonwealth of The
Hispaniola (Fig. 3). The area consists of more than Bahamas became fully independent from Britain in
2,700 oceanic islands, cays (pronounced “keys”), 1973, but currently maintains its membership in the
and rocks that are dispersed among 15 shallow- Commonwealth of Nations. The smaller Turks and
water carbonate banks. Mouchoir, Silver, and Caicos Islands remain part of the British Overseas
Navidad banks in the eastern Turks and Caicos area Territories.
are completely inundated. The banks are separated The majority of the Bahamian Archipelago
by deep-water trenches and strong ocean currents. lies above the Tropic of Cancer (at 23.43ºN), where
These banks and their associated islands extend it is considered part of the subtropics. The climate
between latitudes 20ºN and 28ºN and longitudes is moderated by the Gulf Stream, resulting in mild
71ºW and 80ºW. All of the banks are flooded with temperatures (never freezing) with average highs
marine waters. The combined island land mass and lows of 28.8 °C and 20.8 ºC. Average annual
associated with this immense area comprises only precipitation is 138.9 cm, with the greatest amounts
about 11,000 km2. of rain in summer (May–October) and occasional

Figure 3. Map of the Bahama Islands and the Turks & Caicos Islands.
1
Villa Capulet, Montague Foreshore, P.O. Box N 8893, Nassau, The Bahamas <[email protected]>
2
Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA <[email protected]>
3
Department of Biology, University of Massachusetts Boston, Boston, Massachusetts 02125, USA <[email protected]>
94 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

hurricanes in summer and fall. Surface features and maintained island lists for the Bahamian
are dominated by limestone karst features (eroded Archipelago. Information was gleaned from
limestone surfaces, limestone hills, sinkholes, publications (e.g., Schwartz & Henderson 1988;
and caves). Soils are shallow, mostly alkaline, Franz et al. 1993) and museum indices. Subsequent
and water tables are close to the surface. Coppice updates reflected information that became available
(tropical hardwoods), brackish and salt ponds with from direct observations, communications from
mangroves, maritime marshes, and other coastal researchers in the field, and photographic vouchers
habitats are common throughout the islands. and research reports. The most recent overview of
Pinelands dominate the vegetative communities Bahamian herpetology was Knapp et al. (2011),
on several of the larger islands (Carew & Mylroie which emphasized conservation. In the following
1997). list, we record two additional introduced species:
The herpetofauna of the Bahamian Archipel- Plestiodon sp. on Grand Bahama (Johnson 2011)
ago consists of 48 native species: three amphibians, and Ramphotyphlops braminus on New Providence
30 lizards, 13 snakes, and two freshwater turtles. (A. Davis, pers. comm.). Also, whether an
Also recorded are two crocodilians and fossil turtles introduced species of anole on Grand Bahama is
and tortoises. We have listed more than 1,600 island Anolis carolinensis or A. smaragdinus still has to
occurrences of amphibians and reptiles from nearly be determined (Jonathon Losos, pers. comm.).
250 islands. These occurrences span all 15 banks Island locations were verified using Bahamas
with islands, with the greatest species richness on Land & Surveys 1:25,000 (BLS Series) maps.
the largest islands and banks. Some locations are known or have been known
Tortoises (Chelonoidis alburyorum and other by multiple names and these are indicated. The
endemic species of Chelonoidis) are extinct in the areas of the cays have been estimated using Google
Bahamian Archipelago and the West Indies; Cuban Earth®. Note that coordinates provide only a
Crocodiles (Crocodylus rhombifer) are extirpated general location for some of the smaller cays. We
from the archipelago, but continue to survive in will continue to maintain and update this list as
Cuba; and Bahamian Rock Iguanas (Cyclura spp.) new information is forthcoming. Cays on which
disappeared from the Little Bahamas Bank, but Anolis sagrei and Leiocephalus carinatus have
continue to exist on other Bahamian banks. These been translocated for experimental studies have not
extinction events apparently occurred in the late been identified or listed.
Holocene, but prior to the appearance of humans The most recent overviews of Turks and
(Steadman et al. 2007; Franz & Franz 2009). Caicos herpetology were Reynolds (2011a, 2011b),
Cuba and Hispaniola are the common sources which emphasized conservation. Additional sour-
for most of the Bahamian herpetofauna. None of the ces included Wing and Scudder (1983), Gerber
native species occur naturally in North America. All (1998), Carlson (1999), Newsom and Wing (2004),
of the native species have their affinities with the Bryan et al. (2007), Reynolds and Niemiller (2009,
Neotropics, except for a gecko, which might have 2010a, 2010b), Reynolds (2010, 2012), Reynolds
African ties (e.g., Franz et al. 1996). More than et al. (2010, 2011, 2012), and data provided by G.
20 additional species of frogs, lizards, snakes, and Gerber and M. Welch. Coordinates for the Turks
turtles have been reported as recent introductions, and Caicos were generated by using Google Earth®
possibly gaining entry as released pets or with and verified in Geody®.
horticultural plants from plant nurseries, mostly in
southern Florida. Some have become established. ACKNOWLEDGMENTS
The list of introduced amphibians and reptiles will
continue to expand as more agricultural products For making their indices available at the beginning
are exchanged between the two areas (Lee 2004). of this project we thank the then curators of the
Since 1992, two of us (SDB & RF), using following museums: José Rosado, Museum of
MacLean et al. (1977) as a template, have updated Comparative Zoology (MCZ); the late C. Jack
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 95

McCoy, The Carnegie Museum of Natural History CROOKED-ACKLINS BANK

(CM); Ronald I. Crombie, National Museum of Acklin’s Island (22.40°/-74.00°) (497 km2)
Natural History, Smithsonian Institution (USNM); Osteopilus septentrionalis
John E. Simmons, Natural History Museum, Chelonoidis sp. (F)
Anolis brunneus
University of Kansas (KU); Christopher A. Phillips,
Leiocephalus punctatus
University of Illinois Museum of Natural History- Sphaerodactylus corticola campter
Illinois Natural History Survey UIMNH-INHS, Cubophis vudii raineyi
and all those who have shared their observations, Epicrates chrysogaster schwartzi
including Nancy Albury, Aileen Bainton, Donald Crocodylus sp. (F)
Buden, Eric Carey, Ancellino Davis, Paul Dean, C. Castle Cay (= Castle Island) (22.13°/-74.31°) (1.93 km2)
Kenneth Dodd, Nancy Elliott, Lynn Gape, William Anolis brunneus
Hayes, S. Blair Hedges, Maurice Isaacs, John Leiocephalus punctatus
Iverson, Audrey James, Pat Jodice, Scott Johnson, Sphaerodactylus corticola campter
Charles Knapp, David Knowles, Denis Knowles, Cotton Cay (= Cotton Bay Cay) (22.31°/-74.18°) (0.43 km2)
Kenneth Krysko, David Lee, Jonathan Losos, Anolis brunneus
William Mackin, Melissa Maura, Brian Mealey, South Cay (22.41°/-74.24°) (0.1 km2)
Rick Oliver, Toby Ross, Williams Simmons, Joseph Leiocephalus punctatus ssp.
Wasilewski, and Anthony White. Mark Welch and
Crooked Island (22.73°/-74.20°) (252 km2)
Brian Naqqi Manco reviewed the Turks and Caicos Osteopilus septentrionalis
Island List. Anolis brunneus
Anolis sagrei ordinatus
Leiocephalus punctatus
CAY SAL BANK Sphaerodactylus corticola campter
Cay Sal (23.69°/-80.39°) (1.22 km2) Cubophis vudii raineyi
Anolis fairchildi
Epicrates chrysogaster schwartzi
Anolis sagrei ordinatus
Crocodylus sp. (F)
Anguilla Cays (23.57°/-79.59°) (0.62 km2)
Anolis sagrei ordinatus Bird Rock Cay (22.85°/-74.35°) (0.03 km2)
Leiocephalus punctatus
Cotton Cay (23.49°/-79.51°) (1.45 km2)
Anolis fairchildi Fish Cay (22.49°/-74.25°) (0.88 km2)
Anolis sagrei ordinatus Cyclura rileyi nuchalis
Leiocephalus punctatus
Double Headed Shot Cay (23.93°/-80.47°) (0.20 km2)
Tropidophis curtus curtus Sphaerodactylus corticola campter

Elbow Cay (23.96°/-80.44°) (0.32 km2) Fortune Island (= Long Cay) (22.61°/-74.33°) (24.7 km2)
Anolis sagrei ordinatus Anolis brunneus
Sphaerodactylus nigropunctatus flavicauda Cyclura rileyi nuchalis (PX)
Tropidophis curtus curtus Leiocephalus punctatus
Typhlops biminiensis Goat Cay (22.68°/-74.29°) (0.19 km2)
Leiocephalus punctatus
CONCEPTION BANK
Conception Island (23.83°/-75.12°) (8.17 km2) Guana Cays (four cays: 22.46°/-74.23°; 22.45°/-74.22°;
Osteopilus septentrionalis 22.45°/-74.23°; 22.45°/-74.23°) (0.30 km2)
Anolis sagrei ssp. Leiocephalus punctatus
Anolis smaragdinus Sphaerodactylus corticola campter
Leiocephalus loxogrammus ssp.
North Cay (22.52°/-74.29°) (0.95 km2)
Sphaerodactylus nigropunctatus gibbus
Epicrates striatus ssp. Cyclura rileyi nuchalis
Leiocephalus punctatus
Booby Cay (23.83°/-75.09°) (0.20 km2) Sphaerodactylus corticola campter
Anolis sagrei ssp.
Sphaerodactylus nigropunctatus ssp. Rat Cay (22.67°/-74.28°) (0.06 km2)
Epicrates striatus ssp. Leiocephalus punctatus
96 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

GREAT BAHAMA BANK Alcorine Cay (24.11°/-77.78°) (214 km2)

Andros Island (24.43°/-77.98°) (5,959 km2) Anolis smaragdinus ssp.
Eleutherodactylus rogersi Cyclura cychlura cychlura
Osteopilus septentrionalis Leiocephalus carinatus
Lithobates grylio (I) Cubophis vudii ssp.
Chelonoidis sp. (F) Beach Cay (24.29°/-77.82°) (0.85 km2)
Trachemys terrapen Cyclura cychlura cychlura
Ameiva auberi behringensis
Bigwood Cay (24.37°/-77.75°) (61 km2)
Ameiva auberi kingi Ameiva auberi kingi
Ameiva auberi sideroxylon Leiocephalus carinatus coryi
Ameiva auberi vulturnus
Anolis angusticeps oligaspis Black Creek Cay (24.33°/-77.94°) (0.15 km2)
Anolis distichus distichoides Cyclura cychlura cychlura
Anolis sagrei ordinatus Blue Hole Cay (24.90°/-77.91°) (0.06 km2)
Anolis smaragdinus lerneri Anolis sagrei ssp.
Anolis smaragdinus smaragdinus Booya Cay (Middle Bight) (24.17°/-77.87°) (59 km2)
Cyclura cychlura cychlura Cyclura cychlura cychlura
Leiocephalus carinatus coryi
Calabash Cay (24.90°/-77.92°) (0.02 km2)
Sphaerodactylus copei cataplexus (I)
Anolis distichus ssp.
Sphaerodactylus nigropunctatus flavicauda
Anolis sagrei ssp.
Sphaerodactylus notatus amaurus
Anolis smaragdinus ssp.
Tarentola americana warreni
Sphaerodactylus nigropunctatus ssp.
Cubophis vudii vudii
Sphaerodactylus notatus ssp.
Epicrates striatus fowleri
Tropidophis curtus androsi Fever Cay (Middle Bight) (24.32°/-77.73°) (2.71 km2)
Typhlops biminiensis Anolis distichus distichoides
Typhlops lumbricalis Anolis sagrei ordinatus
Crocodylus sp. (F) Cyclura cychlura cychlura
Leiocephalus carinatus coryi
Mangrove Cay (24.21°/-77.74°) (225 km2)
Gibson Cay (Middle Bight) (24.33°/-77.68°) (0.35 km2)
Eleutherodactylus rogersi
Ameiva auberi kingi
Osteopilus septentrionalis
Anolis sagrei ordinatus
Ameiva auberi kingi x vulturnus
Anolis angusticeps oligaspis Gold Cay (= Billy Island = Bill’s Island) (24.66°/-78.52°)
Anolis distichus distichoides (2.1 km2)
Anolis sagrei ordinatus Anolis sagrei ordinatus
Anolis smaragdinus ssp. High Cay (24.65°/-77.70°) (0.07 km2)
Cyclura cychlura cychlura Anolis sagrei ordinatus
Sphaerodactylus nigropunctatus flavicauda
High Ridge Cay (Middle Bight) (24.28°/-77.78°)
Sphaerodactylus notatus amaurus (5.49 km2)
Cubophis vudii vudii Osteopilus septentrionalis
Epicrates striatus fowleri Anolis sagrei ordinatus
Tropidophis curtus androsi
Hog Cay (South Bight) (24.16°/-77.66°) (4.0 km2)
South Andros (23.95°/-77.67°) (1,160 km2) Osteopilus septentrionalis
Ameiva auberi kingi Anolis smaragdinus ssp.
Anolis angusticeps ssp. Cyclura cychlura cychlura
Anolis distichus distichoides Leiocephalus carinatus ssp.
Anolis sagrei ordinatus
Iguana Island (South Bight) (24.04°/-77.74°) (0.008 km2)
Anolis smaragdinus ssp.
Cyclura cychlura cychlura
Cyclura cychlura cychlura
Leiocephalus carinatus ssp. Linda Cay (= Linder Cay) (24.19°/-77.67°) (15.83 km2)
Sphaerodactylus nigropunctatus ssp. Cyclura cychlura cychlura
Sphaerodactylus notatus ssp. Little Saddleback Cay (24.93°/-77.91°) (0.005 km2)
Cubophis vudii ssp. Sphaerodactylus nigropunctatus ssp.
Typhlops sp. Anolis sagrei ssp.
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 97

Little Sandy Cay (South Bight) (24.10°/-77.68°) (0.037 km2) Sphaerodactylus nigropunctatus flavicauda
Cyclura cychlura cychlura Sphaerodactylus notatus amaurus
Little Wood Cay (Middle Bight) (24.28°/-77.97°) (12.34 km2) Bond Cay (= Bond’s Cay) (25.49°/-77.72°) (2.62 km2)
Cyclura cychlura cychlura Osteopilus septentrionalis
Long Cay (24.74°/-77.77°) (0.03 km2) Ameiva auberi multilineata
Anolis sagrei ssp. Anolis sagrei ordinatus
Leiocephalus carinatus ssp.
Pigeon Cay (24.88°/-77.89°) (0.02 km2) Sphaerodactylus notatus amaurus
Anolis sagrei ordinatus Cubophis vudii vudii
Anolis smaragdinus ssp.
Sphaerodactylus nigropunctatus flavicauda Cat Cay (25.41°/-77.81°) (0.16 km2)
Anolis sagrei ordinatus
Pit Stop Cay (South Bight) (24.04°/-77.73°) (0.05 km2)
Leiocephalus carinatus coryi
Cyclura cychlura cychlura
Leiocephalus carinatus ssp. Chub Cay (25.41°/-77.89°) (1.85 km2)
Ameiva auberi multilineata
Pure Gold (? = Pure Gold Point, South Andros)
Anolis sagrei ordinatus
(23.97°/-77.52)
Sphaerodactylus nigropunctatus flavicauda
Osteopilus septentrionalis
Sphaerodactylus notatus amaurus
Anolis distichus distichoides
Anolis sagrei ordinatus Cubophis vudii vudii
Anolis smaragdinus ssp. Epicrates striatus fowleri

Rat Cay (24.98°/-77.93°) (0.007 km2) Cistern Cay (25.78°/-77.89°) (1.38 km2)
Anolis sagrei ordinatus Osteopilus septentrionalis
Sphaerodactylus notatus amaurus Ameiva auberi multilineata
Anolis sagrei ordinatus
Reid Cay (= Reids Cay, Middle Bight) (24.30°/-77.91°)
(2.57 km2) Devil’s Cay (25.60°/-77.73°) (0.43 km2)
Ameiva auberi ssp. Ameiva auberi multilineata
Leiocephalus carinatus coryi Leiocephalus carinatus coryi
Sphaerodactylus notatus amaurus Frazier’s Hog Cay (= Frazer’s Hog Cay) (25.42°/-77.84°)
Saddleback Cay (24.93°/-77.92°) (0.04 km ) 2 (3.55 km2)
Anolis sagrei ssp. Osteopilus septentrionalis
Anolis smaragdinus ssp. Ameiva auberi multilineata
Leiocephalus carinatus ssp. Anolis angusticeps oligaspis
Sphaerodactylus nigropunctatus ssp. Anolis distichus distichoides
Sandy Cay (South Bight) (24.08°/-77.70°) (2.90 km2) Anolis sagrei ordinatus
Ameiva auberi ssp. Anolis smaragdinus lerneri
Anolis sagrei ordinatus Leiocephalus carinatus coryi
Cyclura cychlura cychlura Sphaerodactylus nigropunctatus flavicauda
Leiocephalus carinatus coryi Sphaerodactylus notatus amaurus
Cubophis vudii vudii Tropidophis curtus curtus
Typhlops biminiensis
Sheep Cay (25.22°/-78.23°) (0.10 km2)
Anolis sagrei ordinatus Goat Cay (25.81°/-77.90°) (0.09 km2)
Anolis sagrei ordinatus
South Joulter Cay (25.29°/-78.12°) (3.94 km2)
Anolis sagrei ordinatus Great Harbour Cay (25.77°/-77.87°) (25.9 km2)
Leiocephalus carinatus coryi Eleutherodactylus rogersi
Osteopilus septentrionalis
Steamer Cay (South Bight) (24.03°/-77.76°) (0.74 km2)
Ameiva auberi multilineata
Cyclura cychlura cychlura
Anolis angusticeps ssp.
William Cay (= Williams Island) (24.66°/-78.46°) (14.69 km2) Anolis distichus distichoides
Leiocephalus carinatus coryi Anolis sagrei ordinatus
Sphaerodactylus notatus amaurus Anolis smaragdinus lerneri
Berry Islands (31 km2) Ctenosaura similis (I)
Alder Cay (25.54°/-77.71°) (0.43 km2) Leiocephalus carinatus coryi
Anolis sagrei ordinatus Sphaerodactylus nigropunctatus flavicauda
98 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Sphaerodactylus notatus amaurus North Bimini (25.75°/-79.28°) (3.74 km2)

Cubophis vudii vudii Ameiva auberi richmondi
Epicrates striatus fowleri Anolis angusticeps oligaspis
Tropidophis curtus ssp. Anolis sagrei ordinatus
Typhlops lumbricalis Anolis smaragdinus lerneri
Alligator mississippiensis (I, PX?) Leiocephalus carinatus coryi
Great Stirrup Cay (25.82°/-77.91°) (1.01 km2) Sphaerodactylus argus (I?)
Anolis sagrei ordinatus Sphaerodactylus notatus amaurus
Anolis smaragdinus smaragdinus Cubophis vudii picticeps
Iguana iguana (I) Epicrates striatus fosteri
Ctenosaura similis (I) Tropidophis curtus curtus
Sphaerodactylus nigropunctatus flavicauda Typhlops biminiensis
Sphaerodactylus notatus amaurus North Cat Cay (= North Cat Island) (25.56°/-79.28°) (1.1 km2)
Hoffman’s Cay (= Holmes Cay) (25.63°/-77.74°) (1.66 km2) Anolis sagrei ordinatus
Ameiva auberi multilineata Anolis smaragdinus lerneri
Anolis sagrei ordinatus South Bimini (25.70°/-79.27°) (8.31 km2)
Anolis smaragdinus lerneri Eleutherodactylus planirostris
Leiocephalus carinatus coryi Osteopilus septentrionalis
Sphaerodactylus nigropunctatus flavicauda Trachemys sp. (I)
Sphaerodactylus notatus amaurus Ameiva auberi richmondi
Lignum Vitae Cay (25.79°/-77.89°) (0.54 km2) Anolis angusticeps oligaspis
Anolis distichus distichoides Anolis distichus biminiensis
Anolis sagrei ordinatus Anolis sagrei ordinatus
Anolis smaragdinus lerneri
Little Harbour Cay (25.58°/-77.72°) (0.94 km2) Leiocephalus carinatus coryi
Ameiva auberi multilineata Sphaerodactylus nigropunctatus flavicauda
Anolis distichus distichoides Sphaerodactylus notatus amaurus
Anolis sagrei ordinatus Cubophis vudii picticeps
Anolis smaragdinus lerneri Epicrates striatus fosteri
Little Stirrup Cay (= Coco Cay) (25.82°/-77.94°) (0.39 km2) Tropidophis curtus curtus
Iguana iguana (I) Typhlops biminiensis
Typhlops lumbricalis
Whale Cay (25.42°/-77.78°) (3.46 km2)
Anolis sagrei ordinatus South Cat Cay (= South Cat Island) (25.53°/-79.26)(0.31 km2)
Ameiva auberi richmondi
Bimini Islands (23 km2) Anolis sagrei ordinatus
East Bimini (25.75°/-79.25°) (5.81 km2) Sphaerodactylus notatus amaurus
Ameiva auberi richmondi
Anolis sagrei ordinatus Tocas Cay (in Bimini Bay) (= Toktos Cay = Sandy Cay)
Leiocephalus carinatus coryi (25.72°/-79.29°) (0.003 km2)
Cubophis vudii picticeps Anolis sagrei ordinatus
Epicrates striatus fosteri Cat Island (24.40°/-75.52°) (389 km2)
Easter Cay (= Easter Island) (not identified) Eleutherodactylus rogersi
Ameiva auberi richmondi Osteopilus septentrionalis
Leiocephalus carinatus coryi Trachemys terrapen (felis)
Cubophis vudii picticeps Ameiva auberi felis
Epicrates striatus fosteri Anolis angusticeps oligaspis
Anolis distichus distichoides x A. d. dapsilis x A. d. ocior
Gun Cay (25.58°/-79.29°) (0.16 km2) Anolis sagrei ordinatus
Ameiva auberi richmondi Anolis smaragdinus smaragdinus
Anolis sagrei ordinatus Leiocephalus carinatus hodsdoni
Anolis smaragdinus lerneri Sphaerodactylus nigropunctatus ssp.
Sphaerodactylus nigropunctatus flavicauda Sphaerodactylus notatus amaurus
Sphaerodactylus notatus amaurus Cubophis vudii vudii
Tropidophis curtus curtus Epicrates striatus ailurus
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 99

Tropidophis curtus barbouri Sphaerodactylus notatus amaurus

Typhlops lumbricalis Cubophis vudii vudii
Alligator Cay (24.54°/-75.64°) (1.0 km2) Tropidophis curtus curtus
Epicrates striatus ailurus Windemere Island (25.07°/-76.12°) (2.69 km2)
Cay Lobos (22.38°/-77.59°) (0.004 km2) Ameiva auberi thoracica
Anolis sagrei ssp. Anolis sagrei ordinatus
Anolis smaragdinus ssp.
Cay Verde (22.03°/-75.20°) (0.16 km2) Cubophis vudii vudii
Ameiva sp.
Anolis sp. Exuma
Leiocephalus carinatus hodsdoni Great Exuma Island (23.50°/-75.77°) (215 km2)
Cubophis sp. Eleutherodactylus rogersi
Osteopilus septentrionalis
Eleuthera (25.20°/-76.24°) (518 km2) Trachemys sp.
Eleutherodactylus rogersi Ameiva auberi obsoleta
Osteopilus septentrionalis Anolis angusticeps oligaspis
Chelonoidis sp. (F) Anolis distichus distichus
Trachemys terrapen Anolis sagrei ordinatus
Ameiva auberi thoracica Anolis smaragdinus smaragdinus
Anolis angusticeps oligaspis Iguana iguana (W)
Anolis distichus ssp. Hemidactylus mabouia (I?)
Anolis sagrei ordinatus Leiocephalus carinatus virescens
Anolis smaragdinus smaragdinus Sphaerodactylus nigropunctatus gibbus
Leiocephalus carinatus virescens Sphaerodactylus notatus amaurus
Hemidactylus mabouia (I?) Cubophis vudii vudii
Sphaerodactylus copei cataplexis (I) Epicrates striatus strigilatus
Sphaerodactylus nigropunctatus nigropunctatus Typhlops lumbricalis
Sphaerodactylus notatus amaurus
Tarentola americana warreni Little Exuma Island (23.43°/-75.59°) (34.58 km2)
Cubophis vudii vudii Eleutherodactylus rogersi
Epicrates striatus strigilatus Osteopilus septentrionalis
Tropidophis curtus barbouri Trachemys sp.
Typhlops lumbricalis Ameiva auberi obsoleta
Anolis distichus distichus
Crocodylus sp. (F)
Anolis sagrei ordinatus
Current Island (25.35°/-76.82°) (8.8 km2) Anolis smaragdinus smaragdinus
Eleutherodactylus rogersi Sphaerodactylus nigropunctatus gibbus
Ameiva auberi thoracica Sphaerodactylus notatus amaurus
Anolis smaragdinus smaragdinus Cubophis vudii vudii
Sphaerodactylus nigropunctatus nigropunctatus Tropidophis curtus barbouri
Sphaerodactylus notatus amaurus Typhlops lumbricalis
Cubophis vudii vudii
Northern Exuma Cays
Harbour Island (25.50°/-76.64°) (2.59 km2) 2nd Cay N of Leaf Cay (24.75°/-76.83°) (0.002 km2)
Anolis distichus ssp. Cyclura cychlura inornata
Anolis sagrei ordinatus
4th Cay N of Leaf Cay (24.76°/-76.83°) (0.006 km2)
Opheodrys aestivus (I)
Anolis sagrei ssp.
North Eleuthera (25.50°/-76.72°) (105.5 km2)
Allen Cay (= Allan’s Cay) (24.75°/-76.84°) (0.065 km2)
Osteopilus septentrionalis
Anolis sagrei ordinatus
Pimlico Cays (= Pimlico Islands) (25.33°/-76.87°) (0.32 km2) Cyclura cychlura inornata
Anolis sagrei ordinatus Sphaerodactylus nigropunctatus
Royal Island (25.52°/-76.84°) (1.71 km2) Cubophis vudii vudii
Eleutherodactylus rogersi Alligator Cay (24.39°/-76.64°) (0.018 km2)
Anolis distichus dapsilis Anolis sagrei ordinatus
Anolis sagrei ordinatus Cyclura cychlura inornata (I)
Anolis smaragdinus smaragdinus Leiocephalus carinatus virescens
100 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Sphaerodactylus nigropunctatus gibbus Leaf Cay (= Allen Cay) (24.75°/-76.84°) (0.044 km2)
Sphaerodactylus notatus amaurus Anolis sagrei ordinatus
Tarentola americana warreni Cyclura cychlura inornata
Cubophis vudii vudii Leiocephalus carinatus virescens
Barn Owl Cay (24.77°/-76.83°) (0.11 km2) Sphaerodactylus n. nigropunctatus x S. n. gibbus
Anolis sagrei ssp. Tarentola americana warreni
Cyclura cychlura inornata Little Bell Island (= Little Bells Cay = Cambridge Cay)
Bell Island (= Bells Cay) (24.30°/-76.56°) (1.41 km2) (24.30°/-76.53°) (1.03 km2)
Eleutherodactylus rogersi Ameiva auberi obsoleta
Osteopilus septentrionalis Anolis sagrei ordinatus
Ameiva auberi obsoleta Sphaerodactylus nigropunctatus gibbus
Anolis distichus distichus Sphaerodactylus notatus amaurus
Anolis sagrei ordinatus
Little Cistern Cay (24.46°/-76.74°) (0.15 km2)
Anolis smaragdinus smaragdinus
Sphaerodactylus notatus amaurus Anolis sagrei ordinatus
Cubophis vudii vudii Sphaerodactylus nigropunctatus gibbus
Epicrates striatus strigilatus Sphaerodactylus notatus amaurus

Bush Hill Cay (24.57°/-76.79°) (0.03 km2) Little Hawksbill Cay (24.46°/-76.76°) (0.16 km2)
Anolis sagrei ordinatus Sphaerodactylus nigropunctatus gibbus
Cyclura rileyi nuchalis (I) Little Norman’s Cay (= Saddle Cay) (24.65°/-76.81)
Sphaerodactylus notatus amaurus (0.22 km2)
Flat Rock Reef Cay (24.76°/-76.83°) (0.044 km2) Anolis sagrei ordinatus
Anolis sagrei ordinatus Leiocephalus carinatus virescens
Cyclura cychlura inornata Sphaerodactylus n. nigropunctatus x S. n. gibbus
Leiocephalus carinatus ssp.
Little Wax Cay (24.56°/-76.79°) (0.19 km2)
Sphaerodactylus sp.
Anolis sagrei ordinatus
Hall’s Pond Cay (24.35°/-76.58°) (1.76 km2)
Long Cay (24.68°/-76.81°) (0.29 km2)
Anolis distichus distichus
Anolis sagrei ordinatus Leiocephalus carinatus virescens
Anolis smaragdinus smaragdinus Long Rock Cay (= West Shroud Cay = Long Cay)
Sphaerodactylus nigropunctatus gibbus (24.40°/-76.66°) (0.12 km2)
Sphaerodactylus notatus amaurus Anolis sagrei ordinatus
Tarentola americana warreni Sphaerodactylus notatus amaurus
Hawksbill Cay (24.48°/-76.77°) (2.76 km2) Tarentola americana warreni
Eleutherodactylus rogersi Cubophis vudii vudii
Ameiva auberi obsoleta Tropidophis curtus ssp.
Anolis distichus distichus Typhlops sp.
Anolis sagrei ordinatus Malabar Cays (24.37°/-76.63°) (0.024 km2)
Anolis smaragdinus smaragdinus
Anolis sagrei ordinatus
Leiocephalus carinatus virescens
Sphaerodactylus nigropunctatus gibbus
Sphaerodactylus notatus amaurus
Tarentola americana warreni Marion Cay (24.37°/-76.60°) (0.022 km2)
Cubophis vudii vudii Anolis sagrei ordinatus
Epicrates striatus strigilatus Sphaerodactylus nigropunctatus gibbus
Tropidophis curtus barbouri
Narrow Water Cay (24.39°/-76.64°) (0.18 km2)
Hawksbill No. 1 Cay (24.46°/-76.76°) (0.017 km ) 2
Ameiva auberi obsoleta
Anolis sagrei ordinatus Anolis sagrei ordinatus
Highborne Cay (= Hyburn Cay) (24.72°/-76.82°) (2.29 km2) Cyclura cychlura inornata (I)
Anolis sagrei ordinatus Leiocephalus carinatus virescens
Sphaerodactylus nigropunctatus gibbus
Hog Cay (24.37°/-76.61°) (0.06 km2)
Sphaerodactylus notatus amaurus
Ameiva auberi obsoleta
Anolis sagrei ordinatus Cubophis vudii vudii
Leiocephalus carinatus virescens Noddy Cay (24.41°/-76.65°) (0.022 km2)
Sphaerodactylus nigropunctatus gibbus Anolis sagrei ordinatus
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 101

Norman’s Cay (24.62°/-76.82°) (3.57 km2) SW Allen’s Cay (= U-Cay = SW Allan’s Cay) (24.74°/-76.84°)
Anolis distichus distichus (0.039 km2)
O’Brien Cay (= O’Brien’s Cay) (24.32°/-76.55°) (0.54 km2) Osteopilus septentrionalis
Anolis sagrei ordinatus Anolis sagrei ordinatus
Leiocephalus carinatus virescens Cyclura cychlura inornata
Sphaerodactylus notatus amaurus Leiocephalus carinatus virescens
Tarentola americana warreni
Pasture Cay (24.32°/-76.56°) (0.043 km2)
Anolis sagrei ordinatus Waderick Wells Cay (= Warderick Wells) (24.39°/-76.62°)
Cyclura cyclura figginsi (I) (1.32 km2)
Sphaerodactylus nigropunctatus gibbus Eleutherodactylus rogersi
Cubophis vudii vudii Ameiva auberi obsoleta
Anolis distichus distichus
Rader’s Rock (24.39°/-76.63°) (0.003 km2) Anolis sagrei ordinatus
Anolis sagrei ordinatus Anolis smaragdinus smaragdinus
Richard’s Rock (24.40°/-76.64°) (0.005 km2) Cyclura cychlura inornata (I, PX)
Anolis sagrei ordinatus Leiocephalus carinatus virescens
Sphaerodactylus nigropunctatus gibbus Sphaerodactylus nigropunctatus gibbus
Tarentola americana warreni Sphaerodactylus notatus amaurus
Tarentola americana warreni
Roberts Cay (24.80°/-76.83°) (0.044 km2)
Cubophis vudii vudii
Cyclura cychlura inornata (I)
Typhlops sp.
Rocky Dundas Cay (24.28°/-76.54°) (0.045 km2)
Wax Cay (24.58°/-76.80°) (0.48 km2)
Anolis sagrei ordinatus
Anolis sagrei ordinatus
Sphaerodactylus notatus amaurus
Anolis smaragdinus smaragdinus
Cubophis vudii vudii
Leiocephalus carinatus virescens
Sandy Cay (24.34°/-76.59°) (0.036 km2)
White Bay Cay (24.34°/76.60°) (0.10 km2)
Anolis sagrei ordinatus
Leiocephalus carinatus virescens
Sphaerodactylus nigropunctatus gibbus
Central Exuma Cays
Ship Channel Cay (24.82°/-76.82°) (1.49 km2)
Big Farmer’s Cay (23.94°/-76.30°) (2.10 km2)
Ameiva auberi focalis
Anolis sagrei ssp.
Anolis sagrei ordinatus
Sphaerodactylus nigropunctatus gibbus
Anolis smaragdinus smaragdinus
Sphaerodactylus notatus amaurus
Leiocephalus carinatus virescens
Sphaerodactylus n. nigropunctatus x S. n. gibbus Bitter Guana Cay (24.14°/-76.42°) (0.88 km2)
Sphaerodactylus notatus amaurus Ameiva auberi obsoleta
Cubophis vudii vudii Anolis sagrei ordinatus
Cyclura cychlura figginsi
Shroud Cay (24.53°/-76.78°) (7.74 km2)
Ameiva auberi obsoleta Bock Cay (= Peace and Plenty Cay) (23.81°/-76.17°) (1.55 km2)
Anolis smaragdinus smaragdinus Ameiva auberi obsoleta
Leiocephalus carinatus virescens Anolis distichus distichus
Sphaerodactylus nigropunctatus gibbus Anolis sagrei ordinatus
Sphaerodactylus notatus amaurus Anolis smaragdinus smaragdinus
Cyclura cychlura figginsi
Soldier Cay (24.33°/-76.56°) (0.24 km2)
Sphaerodactylus nigropunctatus gibbus
Anolis sagrei ordinatus
Tarentola americana warreni
Sooty Cay (24.41°/-76.64°) (0.036 km2)
Cave Cay (23.91°/-76.27°) (1.10 km2)
Anolis sagrei ordinatus
Ameiva auberi obsoleta
South Halls Pond Cay (24.33°/-76.56°) (0.25 km2) Leiocephalus carinatus ssp.
Anolis sagrei ordinatus Sphaerodactylus nigropunctatus gibbus
Southeastern Flat Rock Reef Cay (24.75°/-76.83) Compass Cay (24.27°/-76.52°) (2.87 km2)
(0.007 km2) Eleutherodactylus rogersi
Cyclura cychlura inornata Ameiva auberi obsoleta
Leiocephalus carinatus ssp. Anolis sagrei ordinatus
102 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Anolis smaragdinus smaragdinus Musha Cay (= Moosha Cay) (23.89°/-76.26°) (0.47 km2)
Leiocephalus carinatus virescens Ameiva auberi obsoleta
Sphaerodactylus nigropunctatus gibbus Anolis distichus distichus
Sphaerodactylus notatus amaurus Anolis sagrei ordinatus
Epicrates striatus strigilatus Noddy Cay (= Prickly Pear Cay = Guana Cay = Dildo Cay)
Darby Cay (= Darby Island) (23.85°/-76.22°) (2.18 km2) (off Norman’s Pond Cay) (23.80°/-76.14°) (0.07 km2)
Eleutherodactylus rogersi Cyclura cychlura figginsi
Trachemys sp. (?) Anolis smaragdinus ssp.
Anolis distichus distichus Norman’s Pond Cay (23.78°/-76.13°) (2.18 km2)
Sphaerodactylus nigropunctatus gibbus Anolis sagrei ordinatus
Galliot Cay (= Big Galliot Cay) (23.92°/-76.29°) (0.11 km2) Anolis smaragdinus smaragdinus
Anolis sagrei ordinatus Cyclura cychlura figginsi
Gaulin Cay (= Gaulin Cay South) (24.12°/-76.40°) (0.17 km2) North Adderly Cay (= No Name Cay) (23.79°/-76.12)
Ameiva auberi obsoleta (0.07 km2)
Cyclura cychlura figginsi Ameiva auberi ssp.
Great Guana Cay (24.03°/-76.37°) (12.3 km2) Cyclura cychlura figginsi
Eleutherodactylus rogersi Leiocephalus carinatus virescens
Osteopilus septentrionalis Sphaerodactylus nigropunctatus gibbus
Ameiva auberi obsoleta Pipe Cay (24.24°/-76.52°) (1.33 km2)
Anolis smaragdinus smaragdinus Osteopilus septentrionalis
Cyclura cychlura figginsi (PX) Ameiva auberi obsoleta
Leiocephalus carinatus virescens Anolis sagrei ordinatus
Sphaerodactylus nigropunctatus gibbus Sphaerodactylus notatus amaurus
Sphaerodactylus notatus amaurus Tropidophis curtus barbouri
Tarentola americana warreni Typhlops lumbricalis
Joe Cay (24.25°/-76.50°) (0.53 km2) Rudder Cut Cay (23.88°/-76.24°) (1.63 km2)
Anolis sagrei ordinatus Ameiva auberi obsoleta
Sphaerodactylus notatus amaurus Anolis distichus distichus
Kemp Cay (24.23°/-76.50°) (0.15 km2) Anolis sagrei ordinatus
Anolis sagrei ordinatus Anolis smaragdinus smaragdinus
Leiocephalus carinatus virescens
Leaf Cay (off Norman’s Pond Cay) (23.79°/-76.13°) (0.13 km2) Sphaerodactylus nigropunctatus gibbus
Anolis sagrei ssp. Sphaerodactylus notatus amaurus
Anolis smaragdinus ssp. Tarentola americana warreni
Cyclura cychlura figginsi Cubophis vudii vudii
Lee Stocking Island (23.77°/-76.09°) (2.20 km2) Sampson Cay (24.21°/-76.47°) (2.1 km2)
Osteopilus septentrionalis Ameiva auberi obsoleta
Ameiva auberi obsoleta Anolis distichus distichus
Anolis distichus distichus Anolis sagrei ordinatus
Anolis sagrei ordinatus Anolis smaragdinus smaragdinus
Anolis smaragdinus smaragdinus Sphaerodactylus nigropunctatus gibbus
Cyclura cychlura figginsi (PX) Leiocephalus carinatus ssp.
Tarentola americana warreni
Epicrates striatus strigilatus Staniel Cay (24.17°/-76.44°) (1.69 km2)
Eleutherodactylus rogersi
Lignum Vitae Cay (off Norman’s Pond Cay) (23.80°/-76.15°) Ameiva auberi obsoleta
(0.018 km2) Anolis distichus distichus
Anolis sagrei ordinatus Anolis sagrei ordinatus
Cyclura cychlura figginsi (I) Anolis smaragdinus smaragdinus
Little Farmer’s Cay (23.96°/-76.32°) (0.67 km2) Sphaerodactylus nigropunctatus gibbus
Anolis distichus distichus Sphaerodactylus notatus amaurus
Anolis sagrei ordinatus Tropidophis curtus barbouri
Anolis smaragdinus smaragdinus Typhlops lumbricalis
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 103

Thomas Cay (24.23°/-76.49°) (0.46 km2) Green Cay (= Tongue of Ocean) (24.04°/-77.17°) (2.15 km2)
Leiocephalus carinatus virescens Eleutherodactylus rogersi
White Bay Cay (off Norman’s Pond Cay) (23.80°/-76.15°) Ameiva auberi parvinsulae
(0.055 km2) Anolis distichus distichus
Ameiva auberi obsoleta Anolis sagrei ordinatus
Anolis sagrei ssp. Leiocephalus carinatus virescens
Cyclura cychlura figginsi Sphaerodactylus nigropunctatus gibbus
Leiocephalus carinatus virescens Sphaerodactylus notatus amaurus
Sphaerodactylus nigropunctatus gibbus Cubophis vudii vudii

William’s Cay (23.76°/-76.08°) (0.75 km2) Little San Salvador (= Half Moon Cay) (24.57°/-75.93°)
Anolis sagrei ordinatus (9.26 km2)
Eleutherodactylus rogersi
Southern Exuma Cays Chelonoidis sp. (F)
Barraterre Cay (= Barataria) (23.69°/-76.05°) (11.15 km2) Ameiva auberi thoracica
Anolis distichus distichus Anolis distichus dapsilis
Anolis sagrei ordinatus Anolis sagrei ordinatus
Anolis smaragdinus smaragdinus Anolis smaragdinus smaragdinus
Sphaerodactylus notatus amaurus Cyclura cychlura inornata (I)
Cluff’s Cay (= Clovis Cay) (23.69°/-76.09°) (0.58 km2) Leiocephalus carinatus hodsdoni
Anolis smaragdinus smaragdinus Sphaerodactylus nigropunctatus gibbus
Cyclura cychlura figginsi (I) Sphaerodactylus notatus ssp.
Cubophis vudii vudii
Duck Cay (23.45°/-76.04°) (0.087 km2)
Anolis sagrei ordinatus Goat Cay (24.60°/-75.89°) (0.053 km2)
Anolis sagrei ordinatus
Elizabeth Cay (= Elizabeth Island) (23.51°/-75.73°) (0.42 km2)
Leiocephalus carinatus hodsdoni
Ameiva auberi obsoleta
Leiocephalus carinatus virescens Long Island (23.25°/-75.12°) (596 km2)
Eleutherodactylus rogersi
Guana Cay (= cay SE of Jewfish Cay) (23.44°/-75.91)
Osteopilus septentrionalis
(0.21 km2)
Trachemys sp.
Anolis smaragdinus ssp.
Ameiva auberi obsoleta
Cyclura cychlura figginsi
Anolis angusticeps oligaspis
Sphaerodactylus notatus ssp.
Anolis distichus distichus
Hog Cay (23.39°/-75.48°) (2.91 km2) Anolis sagrei ordinatus
Trachemys sp. Anolis smaragdinus smaragdinus
Sphaerodactylus nigropunctatus gibbus Hemidactylus mabouia (I?)
Jewfish Cay (= Hummingbird Cay) (23.46°/-75.94) Leiocephalus carinatus hodsdoni
(0.96 km2) Sphaerodactylus nigropunctatus gibbus
Anolis sagrei ordinatus Sphaerodactylus notatus amaurus
Sphaerodactylus nigropunctatus gibbus Tarentola americana warreni
Sphaerodactylus notatus amaurus Epicrates striatus strigilatus
Cubophis vudii vudii Cubophis vudii vudii
Tropidophis curtus barbouri
Pigeon Cay (23.47°/-75.62°) (0.088 km2) Typhlops lumbricalis
Ameiva auberi obsoleta
Leiocephalus carinatus virescens Iguana Cay (= Pinder’s Cay) (23.31°/-75.10°) (0.024 km2)
Sphaerodactylus nigropunctatus gibbus Leiocephalus carinatus hodsdoni
Stocking Island (= Stocky Cay?) (23.53°/-75.76°) (2.34 km2) Salt Pond Cay (23.10°/-74.94°) (0.022 km2)
Anolis sagrei ordinatus Leiocephalus carinatus hodsdoni
Anolis smaragdinus smaragdinus Violet Cay (= Long Island)
Sphaerodactylus nigropunctatus gibbus Leiocephalus carinatus hodsdoni
White Cay (= Sandy Cay) (23.40°/-75.44°) (0.14 km2) New Providence Island (25.03°/-77.40°) (207 km2)
Anolis sagrei ssp. Eleutherodactylus planirostris
Cyclura rileyi cristata Gastrophryne carolinensis (I)
Sphaerodactylus nigropunctatus Osteopilus septentrionalis
104 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Lithobates grylio (I) Ragged Island Range

Chelonoidis sp. (F) Great Ragged Island (22.20°/-75.73°) (12.0 km2)
Trachemys decorata (I) Ameiva auberi bilateralis
Trachemys terrapen x T. stejnegeri (I?) Anolis distichus distichus
Ameiva auberi thoracica Anolis sagrei ordinatus
Anolis angusticeps oligaspis Anolis smaragdinus smaragdinus
Anolis distichus distichus Leiocephalus carinatus hodsdoni
Anolis equestris ssp. (I) Sphaerodactylus nigropunctatus porrasi
Anolis smaragdinus smaragdinus Sphaerodactylus notatus amaurus
Anolis sagrei ordinatus Tarentola americana warreni
Basiliscus sp. (W)
Tropidophis curtus barbouri
Cyclura sp. (PE)
Typhlops lumbricalis
Hemidactylus garnotii (I)
Hemidactylus mabouia (I?) Little Ragged Island (22.16°/-75.71°) (2.89 km2)
Leiocephalus carinatus ssp. (F*, I) Ameiva auberi bilateralis
Sphaerodactylus argus (I?) Anolis distichus distichus
Sphaerodactylus copei cataplexis (I) Anolis sagrei ordinatus
Sphaerodactylus nigropunctatus nigropunctatus Anolis smaragdinus smaragdinus
Sphaerodactylus notatus amaurus Leiocephalus carinatus hodsdoni
Cubophis vudii vudii Sphaerodactylus nigropunctatus porrasi
Epicrates striatus strigilatus Sphaerodactylus notatus amaurus
Pantherophis guttatus (I) Tarentola americana warreni
Ramphotyphlops braminus (I) Cubophis vudii vudii
Thamnophis sauritus sackeni (W) Epicrates striatus mccranei
Thamnophis sirtalis sirtalis (W) Typhlops biminiensis
Tropidophis curtus curtus Typhlops lumbricalis
Typhlops biminiensis
Typhlops lumbricalis Ragged Cays
Crocodylus sp. (PE) Hog Cay (22.25°/-75.75°) (2.55 km2)
Athol Island (25.08°/-77.27°) (0.45 km2) Ameiva auberi bilateralis
Eleutherodactylus planirostris Anolis sagrei ordinatus
Anolis sagrei ordinatus Anolis smaragdinus smaragdinus
Anolis smaragdinus smaragdinus Johnson Cay (22.34°/-75.78°) (0.25 km2)
Sphaerodactylus notatus amaurus Leiocephalus carinata hodsdoni
Hog Cay (= Hog Island = Paradise Island) (25.08°/-77.32°) Knife Cay (22.49°/-75.84°) (0.04 km2)
(3.26 km2) Anolis sagrei ordinatus
Osteopilus septentrionalis Leiocephalus carinatus hodsdoni
Ameiva auberi thoracica
Anolis distichus distichus Margaret Cay (22.30°/-75.78°) (1.53 km2)
Anolis equestris (I) Anolis sagrei ssp.
Anolis sagrei ordinatus Epicrates striatus mccraniei
Hemidactylus mabouia (I?) Maycock Cay (22.29°/-75.76°) (1.22 km2)
Cubophis vudii vudii
Ameiva auberi bilateralis
Rose Island (25.10°/-77.16°) (3.33 km2) Sphaerodactylus notatus amaurus
Ameiva auberi thoracica
Nurse Cay (22.47°/-75.84°) (1.95 km2)
Leiocephalus carinatus ssp.
Sphaerodactylus nigropunctatus nigropunctatus Ameiva auberi bilateralis
Epicrates striatus strigilatus Anolis sagrei ordinatus

Salt Cay (25.10°/-77.28°) (0.35 km2) Raccoon Cay (22.37°/-75.81°) (4.54 km2)
Ameiva auberi ssp. Ameiva auberi bilateralis
Anolis sagrei ssp. South Channel Cay (22.50°/-75.85°) (0.07 km2)
Spruce Cay (25.08°/-77.27°) (0.019 km2) Leiocephalus carinatus hodsdoni
Leiocephalus carinatus ssp. Sphaerodactylus notatus amaurus
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 105

Jumentos Cays LITTLE BAHAMA BANK

Flamingo Cay (22.88°/-75.87°) (0.96 km2) Abaco
Ameiva auberi bilateralis Great Abaco (26.38°/-77.12°) (1,681 km2)
Anolis distichus distichus Eleutherodactylus planirostris
Anolis sagrei ordinatus Osteopilus septentrionalis
Anolis smaragdinus smaragdinus Chelonoidis alburyorum (F)
Leiocephalus carinatus hodsdoni Trachemys sp. (I?)
Sphaerodactylus nigropunctatus porrasi Anolis distichus ssp. (I?)
Sphaerodactylus notatus amaurus Anolis smaragdinus (I)
Tarentola americana warreni Anolis sagrei ordinatus (F*)
Cubophis vudii ssp. Cyclura sp. (F)
Pear Cay (23.13°/-75.53°) (0.03 km2) Leiocephalus carinatus armouri
Anolis sagrei ordinatus Sphaerodactylus notatus peltastes
Leiocephalus carinatus hodsdoni Cubophis sp. (F)
Sphaerodactylus nigropunctatus porrasi Cubophis vudii aterrimus
Water Cay (23.01°/-75.72°) (0.51 km2) Epicrates exsul
Ameiva auberi bilateralis Epicrates striatus (F)
Anolis distichus distichus Pantherophis alleghaniensis ssp. (I)
Anolis sagrei ordinatus Typhlops lumbricalis
Anolis smaragdinus smaragdinus Typhlops sp. (F)
Leiocephalus carinatus hodsdoni Crocodylus rhombifer (F)
Sphaerodactylus nigropunctatus porrasi Little Abaco Island (26.89°/-77.64°) (60 km2)
Sphaerodactylus notatus amaurus Eleutherodactylus planirostris
Tarentola americana warreni Osteopilus septentrionalis
Anolis sagrei ordinatus
GREAT INAGUA BANK Leiocephalus carinatus armouri
Great Inagua (21.01°/-73.18°) (1,552 km2) Sphaerodactylus notatus peltastes
Eleutherodactylus planirostris (I?)
Cubophis vudii aterrimus
Osteopilus septentrionalis (I?)
Epicrates exsul
Trachemys stejnegeri malonei (I?)
Typhlops lumbricalis
Ameiva maynardii maynardii
Ameiva maynardii uniformis Abaco Cays
Anolis scriptus leucophaeus Cave Cay (26.87°/-77.90°) (1.76 km2)
Aristelliger barbouri Anolis sagrei ordinatus
Leiocephalus inaguae Leiocephalus carinatus armouri
Sphaerodactylus inaguae Crab Cay (26.81°/-77.36°) (0.81 km2)
Sphaerodactylus notatus atactis (I?) Anolis sagrei ordinatus
Cubophis vudii utowanae Leiocephalus carinatus armouri
Epicrates chrysogaster relicquus Sphaerodactylus notatus peltastes
Tropidophis canus
Typhlops paradoxus Elbow Cay (26.52°/-76.97°) (4.51 km2)
Crocodylus sp. (W) Osteopilus septentrionalis
Anolis sagrei ordinatus
Sheep Island (= Sheep Cay) (21.13°/-73.59°) (0.25 km2)
Leiocephalus carinatus armouri
Ameiva maynardii maynardii
Sphaerodactylus notatus peltastes
Aristelliger barbouri
Epicrates exsul
Anolis scriptus leucophaeus
Sphaerodactylus inaguae Fiddle Cay (26.81°/-77.35°) (0.088 km2)
Cubophis vudii utowanae Leiocephalus carinatus armouri
Epicrates chrysogaster relicquus Sphaerodactylus notatus peltastes
Fox’s Island (not identified)
LITTLE INAGUA BANK
Anolis sagrei ordinatus
Little Inagua (21.50°/-73.00°) (127 km2)
Ameiva maynardii parvinaguae Great Guana Cay (26.67°/-77.11°) (4.83 km2)
Aristelliger barbouri Anolis sagrei ordinatus
Anolis scriptus leucophaeus Leiocephalus carinatus armouri
Hypsirhynchus parvifrons (W?) Sphaerodactylus notatus peltastes
106 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Green Turtle Cay (26.77°/-77.33°) (4.16 km2) Trachemys sp. (I)

Anolis sagrei ordinatus Anolis carolinensis or A. smaragdinus (I)
Leiocephalus carinatus armouri Anolis distichus ssp. (I?)
Sphaerodactylus notatus peltastes Anolis equestris (I)
Epicrates exsul Anolis sagrei ordinatus
Joe’s Cay (27.10°/-78.06°) (0.137 km2) Leiocephalus carinatus armouri
Leiocephalus carinatus armouri Plestiodon sp. (I)
Sphaerodactylus notatus peltastes
Little Sale Cay (27.04°/-78.17°) (0.096 km2) Cubophis vudii aterrimus
Leiocephalus carinatus armouri Pantherophis guttatus (I)
Mangrove Cay (26.87°/-77.72°) (0.85 km2) Epicrates exsul
Anolis sagrei ordinatus Typhlops lumbricalis
Sphaerodactylus notatus peltastes Storeria dekayi (I)
Crocodylus acutus (PE)
Manjack Cay (= Nunjack Cay) (26.83°/-77.37°) (2.70 km2)
Anolis sagrei ordinatus Burrows Cay (26.43°/-77.78°) (0.38 km2)
Leiocephalus carinatus armouri Anolis sagrei ordinatus
Sphaerodactylus notatus peltastes Leiocephalus carinatus armouri
Man O’War Cay (26.60°/-77.00°) (1.12 km2) Sweeting’s Cay (26.62°/-77.87°) (9.33 km2)
Anolis sagrei ordinatus Osteopilus septentrionalis
Hemidactylus garnotii (I) Anolis sagrei ordinatus
Moraine Cay (27.04°/-77.77°) (0.13 km2) Water Cay (26.44°/-77.77°) (0.18 km2)
Anolis sagrei ordinatus Anolis sagrei ordinatus
Leiocephalus carinatus armouri
Mores Island (= Moore’s Island) (26.30°/-77.56°) (16.4 km2)
Sphaerodactylus notatus peltastes
Chelonoidis sp. (F)
Typhlops lumbricalis
Anolis sagrei ordinatus
Leiocephalus carinatus armouri Wood Cay (26.74°/-79.01°) (0.05 km2)
Sphaerodactylus notatus peltastes Anolis sagrei ordinatus
Leiocephalus carinatus armouri
Pensacola Cays (26.98°/-77.67°) (1.94 km2)
Sphaerodactylus notatus peltastes
Osteopilus septentrionalis
Anolis sagrei ordinatus
Leiocephalus carinatus armouri
MAYAGUANA BANK
Mayaguana Island (22.37°/-72.93°) (285 km2)
Sphaerodactylus notatus peltastes
Chelonoidis sp. (F)
Powell Cay (26.91°/-77.48°) (1.21 km2) Anolis scriptus mariguanae
Anolis sagrei ordinatus Sphaerodactylus mariguanae
Leiocephalus carinatus armouri Crocodylus sp. (F)
Rocky Harbour Cay (26.81°/-77.51°) (1.45 km2) Booby Cay (22.32°/-72.72°) (0.51 km2)
Anolis sagrei ordinatus Anolis scriptus mariguanae
Leiocephalus carinatus armouri Cyclura carinata bartschi
Stranger Cay (= Strangers Cay) (27.12°/-78.08°) (1.41 km2) Sphaerodactylus mariguanae
Osteopilus septentrionalis
Anolis sagrei ordinatus MIRA POR VOS BANK
Leiocephalus carinatus armouri South Cay (22.09°/-74.53°) (0.58 km2)
Sphaerodactylus notatus peltastes Leiocephalus punctatus ssp.

Grand Bahama Island (26.65°/-78.50°) (1,373 km2) RUM CAY BANK

Eleutherodactylus planirostris Rum Cay (23.67°/-74.87°) (78 km2)
Gastrophryne carolinensis (I) Osteopilus septentrionalis
Hyla squirella (I) Anolis distichus ocior
Osteopilus septentrionalis Anolis sagrei ordinatus
Lithobates clamitans (W) Leiocephalus loxogrammus loxogrammus
Lithobates sphenocephalus (I) Sphaerodactylus corticola corticola
Terrapene carolina (W) Sphaerodactylus nigropunctatus decoratus
Trachemys scripta elegans (I) Sphaerodactylus notatus amaurus
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 107

PLANA CAYS BANK Goulding Cay (24.05°/-74.43°) (0.02 km2)

East Plana Cay (22.61°/-73.52°) (7.0 km2) Cyclura rileyi rileyi
Anolis brunneus Green Cay (24.14°/-74.51°) (0.04 km2)
Anolis scriptus ssp. Anolis distichus ocior
Leiocephalus greenwayi Cyclura rileyi rileyi
Sphaerodactylus corticola apporox Sphaerodactylus corticola soter
West Plana Cay (22.59°/-73.61°) (9.0 km2) High Cay (23.95°/-74.48°) (0.13 km2)
Anolis brunneus Cyclura rileyi rileyi (PX)
Anolis scriptus sularum
Sphaerodactylus corticola apporox Little Green Cay (= Gauntlet Cay = Gaulin Cay = Hawksnest
Cay = Goulding Cay (24.14°/-74.47°) (0.011 km2)
SAMANA BANK Cyclura rileyi rileyi (PX)
Samana Cay (= Atwood’s Cay) (23.08°/-73.75°) (39 km2) Epictia columbi
Anolis scriptus sularum Low Cay (23.93°/-74.48°) (0.10 km2)
Leiocephalus punctatus picinus Cyclura rileyi rileyi
Sphaerodactylus corticola ssp. Sphaerodactylus corticola soter
Small Cay (= “Propeller” Cay) (off Samana Cay) Man Head Cay (24.12°/-74.45°) (0.024 km2)
(23.06°/-73.74°) (0.08 km2) Anolis distichus ocior
Anolis scriptus sularum Cyclura rileyi rileyi
West Booby Cay (23.09°/-73.65°) (0.72 km2) Sphaerodactylus corticola soter
Anolis scriptus sularum Pokus Cay (23.94°/-74.48°) (0.018 km2)
Cyclura rileyi rileyi (PX)
SAN SALVADOR BANK White Cay (in Grahams Harbour) (24.16°/-74.48°) (0.019 km2)
San Salvador Island (= Watling’s Island) (24.00°/-74.50°)
Cyclura rileyi rileyi (PX)
(163 km2)
Eleutherodactylus rogersi CAICOS BANK
Hyla squirella (W) West Caicos (21.65°/-72.47°) (22.8 km2)
Osteopilus septentrionalis Anolis scriptus scriptus
Chelonoidis sp. (F) Cyclura carinata (PX)
Trachemys sp. (PX) Leiocephalus psammodromus
Anolis distichus ocior Sphaerodactylus caicosensis
Anolis sagrei ordinatus Spondylurus caicosae
Cyclura rileyi rileyi
Hemidactylus mabouia (I?) French Cay (21.51°/-72.20°) (0.13 km2)
Leiocephalus loxogrammus parnelli Anolis scriptus scriptus
Sphaerodactylus corticola soter Aristelliger hechti
Epictia columbi Cyclura carinata
Crocodylus sp. (F) Providenciales (21.78°/-72.28°) (122.2 km2)
Barn Cay (in Great Lake) (24.08°/-74.50°) (0.17 km2) Eleutherodactylus planirostris (I)
Cyclura rileyi rileyi (PX) Osteopilus septentrionalis (I)
Rhinella marina (W)
Guana Cay (in Little Hermitage Lake) (24.09°/-74.48) Trachemys scripta elegans (W)
(0.011 km2) Anolis equestris ssp. (W)
Cyclura rileyi rileyi Anolis scriptus scriptus
Pidgeon Cay (= Pigeon Cay) (in Great Lake) (24.02°/-74.50°) Cyclura carinata (in Chalk Sound)
(0.071 km2) Hemidactylus mabouia (I?)
Cyclura rileyi rileyi Iguana iguana (W)
Leiocephalus psammodromus
San Salvador Cays
Sceloporus sp. (W)
Cato Cay (= Catto Cay) (24.14°/-74.47°) (0.013 km2) Sphaerodactylus caicosensis
Cyclura rileyi rileyi (PX) Spondylurus caicosae
Sphaerodactylus corticola soter Varanus exanthematicus (W)
Cut Cay (in Rice Bay) (24.13°/-74.46°) (0.028 km2) Epicrates chrysogaster chrysogaster
Cyclura rileyi rileyi (I) Ramphotyphlops braminus (I)
108 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Tropidophis greenwayi lanthanus Leiocephalus psammodromus

Typhlops c.f. platycephalus Sphaerodactylus caicosensis
Uromacer catesbyi or Opheodrys aestivus (ID unclear) (W) Spondylurus caicosae
Caicos Cays North Caicos (21.93°/-71.98°) (116 km2)
Dellis Cay (21.89°/-72.07°) (2.10 km2) Eleutherodactylus planirostris (I)
Anolis scriptus scriptus Osteopilus septentrionalis (I)
Cyclura carinata (tail drags) Anolis scriptus scriptus
Leiocephalus psammodromus Aristelliger hechti
Sphaerodactylus caicosensis Cyclura carinata (W)
Epicrates chrysogaster chrysogaster Hemidactylus mabouia (I?)
Leiocephalus psammodromus
Donna Cay (21.83°/-72.13°) (0.36 km2) Sphaerodactylus caicosensis
Anolis scriptus scriptus Spondylurus caicosae
Cyclura carinata Epicrates chrysogaster chrysogaster
Fort George Cay (21.89°/-72.09°) (0.85 km2) Tropidophis greenwayi lanthanus
Anolis scriptus scriptus Typhlops c.f. platycephalus
Cyclura carinata (PX) Bay Cay (21.93°/-71.89)
Leiocephalus psammodromus Anolis scriptus scriptus
Sphaerodactylus caicosensis Sphaerodactylus caicosensis
Little Water Cay (21.83°/-72.15°) (0.6 km2) Spondylurus caicosae
Anolis scriptus scriptus East Bay (21.89°/-71.88)
Cyclura carinata Anolis scriptus scriptus
Leiocephalus psammodromus Cyclura carinata
Sphaerodactylus caicosensis
Major Hill Cay (21.94°/-71.91)
Lizard, Bird, And Sinking Cays (21.84°/-72.13) Cyclura carinata
Anolis scriptus scriptus
Cyclura carinata Middle Caicos (= Grand Caicos Island) (21.78°/-71.72)
(144 km2)
Mangrove Cay (21.82°/-72.14°) (1.14 km2) Eleutherodactylus planirostris (I)
Anolis scriptus scriptus Osteopilus septentrionalis (I)
Cyclura carinata Chelonoidis sp. (F)
Parrot Cay (21.92°/-72.06W°) (5.6 km2) Anolis scriptus scriptus
Osteopilus septentrionalis (I) Cyclura carinata (PX)
Anolis scriptus scriptus Hemidactylus mabouia (I?)
Hemidactylus mabouia (I?) Leiocephalus psammodromus
Leiocephalus psammodromus Sphaerodactylus caicosensis
Sphaerodactylus caicosensis Spondylurus caicosae
Spondylurus caicosae Epicrates chrysogaster chrysogaster
Epicrates chrysogaster chrysogaster Tropidophis greenwayi lanthanus
Typhlops c.f. platycephalus
Pine Cay (21.87°/-72.09°) (4.21 km2)
Osteopilus septentrionalis (I) Iguana Cay (between Middle and East Caicos)
Trachemys stejnegeri malonei (I, E?) (21.76°/-71.60°) (0.06 km2)
Anolis scriptus scriptus Cyclura carinata
Cyclura carinata Joe Grant’s Cay (between Middle and East Caicos)
Hemidactylus mabouia (I?) (21.76°/-71.61°) (5.1 km2)
Leiocephalus psammodromus Cyclura carinata (tail drags)
Sphaerodactylus caicosensis Epicrates chrysogaster chrysogaster
Spondylurus caicosae
Pelican Cay (21.83°/-71.72) (< 0.1 km2)
Stubbs Cay (21.88°/-72.08) (~0.9 km2) Anolis scriptus scriptus
Anolis scriptus scriptus Sphaerodactylus caicosensis
Leiocephalus psammodromus
East Caicos (21.71°/-71.51°) (90 km2)
Water Cay (21.85°/-72.12°) (3.5 km2) Anolis scriptus scriptus
Anolis scriptus scriptus Aristelliger hechti
Cyclura carinata Cyclura carinata
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 109

Leiocephalus psammodromus Sphaerodactylus caicosensis

Sphaerodactylus caicosensis Spondylurus caicosae
Spondylurus caicosae Epicrates chrysogaster chrysogaster
Epicrates chrysogaster chrysogaster
Long Cay (S of South Caicos) (21.47°/-71.56°) (1.0 km2)
Tropidophis greenwayi lanthanus
Anolis scriptus scriptus
Typhlops c.f. platycephalus
Cyclura carinata (I)
Horse Cay (21.53°/-71.51°) (0.01 km2) Leiocephalus psammodromus
Cyclura carinata Sphaerodactylus caicosensis
J.A.G.S. Mccartney Cay (21.62°/-71.49°) (11.4 km2) Spondylurus caicosae
Cyclura carinata Epicrates chrysogaster chrysogaster (PX)
Tropidophis greenwayi lanthanus
Middle Creek Cay (21.60°/-71.49°) (0.47 km2)
Cyclura carinata Middleton Cay (21.48°/-71.58°) (0.05 km2)
Cyclura carinata
Plandon Cay (21.58°/-71.49°) (0.22 km2) Sphaerodactylus caicosensis
Cyclura carinata Spondylurus caicosae
Sail Rock (21.58°/-71.52) Tropidophis greenwayi lanthanus
Cyclura carinata Six Hills Cay (includes East and West cays) (21.46°/-71.63°)
South Caicos (= Turks Island in late 19th Century) (0.09 km2)
(21.52°/-71.50°) (21.2 km2) Anolis scriptus scriptus
Anolis scriptus scriptus Aristelliger hechti
Cyclura carinata (PX) Cyclura carinata
Hemidactylus mabouia (I?) Sphaerodactylus caicosensis
Sphaerodactylus caicosensis Spondylurus caicosae
Spondylurus caicosae White Cay (21.18°/-71.80°) (0.03 km2)
Epicrates chrysogaster chrysogaster (PX) Cyclura carinata
Tropidophis greenwayi lanthanus
Big Ambergris Cay (21.29°/-71.63°) (4.28 km2) TURKS BANK
Eleutherodactylus planirostris (I) Grand Turk (= Turks Island) (21.46°/-71.13°) (17.39km2)
Osteopilus septentrionalis (W) Chelonoidis sp. (F)
Anolis scriptus scriptus Anolis scriptus scriptus
Aristelliger hechti Cyclura carinata (PX, W)
Cyclura carinata Hemidactylus mabouia (I?)
Hemidactylus mabouia (I) Iguana iguana (W)
Leiocephalus psammodromus Leiocephalus psammodromus (PX)
Sphaerodactylus caicosensis Sphaerodactylus mariguanae (I, PX?)
Spondylurus caicosae Sphaerodactylus underwoodi
Spondylurus turksae (PX)
Epicrates chrysogaster chrysogaster
Epicrates chrysogaster chrysogaster (PX)
Tropidophis greenwayi greenwayi
Pantherophis guttatus (W)
Typhlops c.f. platycephalus
Ramphotyphlops braminus (I)
Bush Cay (21.20°/-71.63°) (0.08 km2)
Big Sand Cay (21.19°/-71.25°) (0.58 km2)
Anolis scriptus scriptus
Anolis scriptus scriptus
Cyclura carinata (I)
Cyclura carinata
Fish Cays (21.37°/-71.62°) (0.1 km2) Leiocephalus psammodromus
Anolis scriptus scriptus Sphaerodactylus underwoodi
Cyclura carinata
Cotton Cay (21.36°/-71.15°) (1.13 km2)
Indian Cay (21.19°/-71.79°) (0.05 km2) Anolis scriptus scriptus
Anolis scriptus scriptus Cyclura carinata (PX)
Cyclura carinata Leiocephalus psammodromus (PX)
Little Ambergris Cay (21.30°/-71.69°) (6.59 km2) Spondylurus turksae
Anolis scriptus scriptus East Cay (= Martin Pinzon) (21.36°/-71.09°) (0.46 km2)
Aristelliger hechti Anolis scriptus scriptus
Cyclura carinata Leiocephalus psammodromus
Leiocephalus psammodromus Sphaerodactylus underwoodi
110 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Gibbs Cay (21.44°/-71.11°) (0.06 km2)

Anolis scriptus scriptus
Leiocephalus psammodromus
Sphaerodactylus underwoodi
Spondylurus turksae
Epicrates chrysogaster chrysogaster
Long Cay (21.41°/-71.10°) (0.23 km2)
Anolis scriptus scriptus
Cyclura carinata
Leiocephalus psammodromus
Sphaerodactylus underwoodi
Pear Cay (21.37°/-71.09°) (0.11 km2)
Leiocephalus psammodromus
Sphaerodactylus underwoodi
Typhlops c.f. platycephalus
Penniston Cay (21.38°/-71.12°) (0.03 km2)
Leiocephalus psammodromus
Round Cay (21.44°/-71.11°) (0.01 km2)
Leiocephalus psammodromus
Salt Cay (21.32°/-71.20°) (6.74 km2)
Anolis scriptus scriptus
Cyclura carinata
Hemidactylus mabouia (I?)
Leiocephalus psammodromus (PX)
Sphaerodactylus underwoodi
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 111

CAYMAN ISLANDS

Arthur C. Echternacht1
The Cayman Islands consist of four islands that Anolis garmani (W)
support fully terrestrial vegetation: Grand Cayman Anolis sagrei (I)
Aristelliger praesignis praesignis
(maximum elevation 18 m), Cayman Brac (43 Cyclura lewisi
m), Little Cayman (12 m), and Owen Island (~2 Cyclura caymanensis (W?)
m). Owen Island is located inside the barrier Gonatodes albogularis (W)
reef at the western edge of South Hole Sound Hemidactylus mabouia (I?)
off the south coast of Little Cayman, from which Iguana iguana (I)
Leiocephalus carinatus varius
it is separated by about 200 m of shallow water.
Ophisaurus ventralis (W)
Several small mangrove islands along the western Sphaerodactylus argivus lewisi
and southwestern edges of North Sound on Grand Cubophis caymanus
Cayman are separated from one another or the Diadophis punctatus (W)
extensive mainland mangrove forest by deep, Pantherophis guttatus (I)
Ramphotyphlops braminus (I)
narrow channels. Because the substrate in which the
Tretanorhinus variabilis lewisi
island mangroves are rooted is entirely submerged Tropidophis caymanensis
at high tide (F. J. Burton, pers. comm.) and faunal Typhlops caymanensis
surveys have not been conducted on these islands, Crocodylus acutus (PX, W, F*)
they are not included in the checklist. Crocodylus acutus x C. rhombifer (W)
Crocodylus rhombifer (F*, E)
Twenty-two native species of amphibians
and reptiles have been documented for the Cayman LITTLE CAYMAN BANK
Islands. One of these, Crocodylus rhombifer, Little Cayman (19.69°/-80.05°) (28 km2)
is extinct. Trachemys decussata angusta is not Osteopilus septentrionalis
included in the total because it is not known Anolis maynardii
whether it is native or introduced. An additional 11 Anolis sagrei sagrei
Aristelliger praesignis praesignis
non-native (introduced) species have been reported
Celestus maculatus
on Grand Cayman and some of these have become Cyclura caymanensis
established. In a few cases, species native to one Leiocephalus carinatus granti (I?)
of the islands have been introduced on another. Sphaerodactylus argivus bartschi
The herpetofauna of the Caymans was reviewed in Cubophis ruttyi
Tropidophis parkeri
detail by Grant (1940) and updated by Seidel and
Crocodylus acutus (PX)
Franz (1994) and Echternacht et al. (2011).
Owen Island (19.66°/-80.06°) (< 0.005 km2)
GRAND CAYMAN BANK Aristelliger praesignis praesignis
Grand Cayman (19.32°/-81.26°) (197 km2) Sphaerodactylus argivus bartschi
Rhinella marina (W)
Eleutherodactylus planirostris CAYMAN BRAC BANK
Gastrophryne carolinensis (I) Cayman Brac (19.72°/-79.80°) (38 km2)
Osteopilus septentrionalis Eleutherodactylus planirostris
Trachemys decussata angusta (I?) Osteopilus septentrionalis
Trachemys scripta elegans (I) Anolis conspersus lewisi
Anolis carolinensis (W) Anolis maynardii (I)
Anolis conspersus conspersus Anolis luteosignifer
Anolis equestris ssp. (W?) Aristelliger praesignis praesignis

1
Department of Ecology & Evolutionary Biology, University of Tennessee, Knoxville, Tennessee 37996 <[email protected]>
112 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Celestus maculatus Cubophis fuscicauda

Cyclura caymanensis Tropidophis schwartzi
Leiocephalus carinatus granti Typhlops epactius
Sphaerodactylus argivus argivus Crocodylus acutus (PX)

Figure 4. Map of the Cayman Islands. See Figure 5 for location of these islands relative to Cuba.
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 113

THE CUBAN ARCHIPELAGO

Alberto R. Estrada1

The Cuban Archipelago, with a total area of of Cuban researchers Luis M. Díaz and Antonio
approximately 110,900 km², contains the largest Cádiz has shed new light on the amphibians of
of the West Indian islands. In addition to the main Cuba (Díaz & Cádiz, 2008).
island, the archipelago includes the Isle of Pines
(Isla de la Juventud) and nearly 1,500 islets or ACKNOWLEDGEMENTS
keys (Fig. 5). Located at the entrance to the Gulf Many people have contributed in one way or
of Mexico, the entire archipelago is home to a another to this effort, for which I am grateful.
rich fauna of over 220 species of amphibians and In alphabetical order: Roberto Alonso, Giraldo
reptiles. Alayón, Rafael Alayo, Luis F. de Armas, Jorge de
The first major compilation of the Cuban la Cruz, Luis M. Díaz, Orlando H. Garrido, Víctor
herpetofauna appeared in volumes IV and VIII of González, John Guarnaccia, S. Blair Hedges,
the overview by Ramón de la Sagra (1838–1861) Robert W. Henderson, Martjan Lammertink,
of the physical, political, and natural history of Alejandro Llanes, Luis V. Moreno, Antonio A.
Cuba. The herpetological sections, written by Jean- Mignucci, Julio Novo Rodríguez, Carlos Peña,
Théodore Cocteau and Gabriel Bibron, listed 34 Antonio Pérez-Asso, Antonio Perera, Robert
species, of which 14 were new. Powell, Celia A. Puerta, Alfredo Rams, Rubén
Since then, numerous contributions have Regalado, Ariel Rodríguez, Roberto Rodríguez
documented the presence of frogs and reptiles in Soberón, Gilberto Silva, Eduardo Solana, Richard
Cuba. Among the most important are Gundlach Thomas, and Alejandro Torres.
(1867, 1880), Barbour and Ramsden (1919), Alayo
(1955), Jaume (1966), Buide (1967), Garrido CUBA BANK
(1973a, 1973b, 1976, 1980), Garrido and Jaume Isla de Cuba (22.00°/-78.90°) (105,806 km2)
Eleutherodactylus acmonis
(1984), Garrido et al. (1986), Estrada (1992, 1993a, Eleutherodactylus adelus
1993b, 1994), Estrada and Novo (1984), Estrada Eleutherodactylus albipes
and Rodríguez (1985), Estrada et al. (1987), and Eleutherodactylus atkinsi atkinsi
Novo et al. (1987). Many of these works addressed Eleutherodactylus atkinsi estradai
specific regions of the Cuban Archipelago, Eleutherodactylus auriculatus
Eleutherodactylus bartonsmithi
including many of the small satellite islands and Eleutherodactylus blairhedgesi
archipelagos off the northern and southern coasts Eleutherodactylus bresslerae
of the main island. Eleutherodactylus casparii
Among Cuban herpetologists, Orlando H. Eleutherodactylus cubanus
Eleutherodactylus cuneatus
Garrido stands out, both for his descriptions of
Eleutherodactylus dimidiatus
many species and his surveys of many satellite Eleutherodactylus eileenae
islands. Albert Schwartz, often in collaboration Eleutherodactylus emiliae
with Garrido, also made substantive contributions Eleutherodactylus erythroproctus
to Cuban herpetology. More recently, S. Blair Eleutherodactylus etheridgei
Eleutherodactylus feichtingeri
Hedges, collaborating with Richard Thomas and Eleutherodactylus glamyrus
several Cuban scientists, has described a number Eleutherodactylus goini
of new species. During the past few years, the work Eleutherodactylus greyi

1
Producciones Eleuth Inc., PMB 256-35 J. C. de Borbon, Ste. 67, Guaynabo, Puerto Rico 00969-5375, USA <[email protected]>
114 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Eleutherodactylus guanahacabibes Eleutherodactylus thomasi zayasi

Eleutherodactylus guantanamera Eleutherodactylus toa
Eleutherodactylus gundlachi Eleutherodactylus tonyi
Eleutherodactylus iberia Eleutherodactylus turquinensis
Eleutherodactylus intermedius Eleutherodactylus varians
Eleutherodactylus ionthus Eleutherodactylus varleyi
Eleutherodactylus jaumei Eleutherodactylus zeus
Eleutherodactylus klinikowskii Eleutherodactylus zugi
Eleutherodactylus leberi Osteopilus septentrionalis
Eleutherodactylus limbatus Lithobates catesbeianus (I)
Eleutherodactylus maestrensis Peltophryne cataulaciceps
Eleutherodactylus mariposa Peltophryne empusa
Eleutherodactylus melacara Peltophryne florentinoi
Eleutherodactylus michaelschmidi Peltophryne fustiger
Eleutherodactylus olibrus Peltophryne gundlachi
Eleutherodactylus orientalis Peltophryne longinasus loginasus
Eleutherodactylus pezopetrus Peltophryne longinasus cajalbanensis
Eleutherodactylus pinarensis Peltophryne longinasus dunni
Eleutherodactylus planirostris Peltophryne longinasus ramsdeni
Eleutherodactylus principalis Peltophryne peltocephala
Eleutherodactylus ricordii Peltophryne taladai
Eleutherodactylus riparius Rhinella marina (I, E)
Eleutherodactylus rivularis Chelonoidis cubensis (F)
Eleutherodactylus ronaldi Trachemys decussata decussata
Eleutherodactylus simulans Trachemys decussata angusta
Eleutherodactylus symingtoni Ameiva auberi auberi
Eleutherodactylus tetajulia Ameiva auberi abducta
Eleutherodactylus thomasi thomasi Ameiva auberi atrothorax
Eleutherodactylus thomasi trinidadensis Ameiva auberi cacuminis

Figure 5. Map of Cuba and surrounding islands.

POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 115

Ameiva auberi citra Anolis guazuma

Ameiva auberi denticola Anolis homolechis
Ameiva auberi garridoi Anolis imias
Ameiva auberi gemea Anolis incredulus
Ameiva auberi granti Anolis inexpectatus
Ameiva auberi hardyi Anolis isolepis
Ameiva auberi llanensis Anolis juangundlachi
Ameiva auberi orlandoi Anolis jubar jubar
Ameiva auberi paulsoni Anolis jubar albertschwartzi
Ameiva auberi peradusta Anolis jubar cocoensis
Ameiva auberi procer Anolis jubar cuneus
Ameiva auberi pullata Anolis jubar gibarensis
Ameiva auberi sabulicolor Anolis jubar maisensis
Ameiva auberi schwartzi Anolis jubar oriens
Ameiva auberi ustulata Anolis jubar yaguajayensis
Ameiva auberi zugi Anolis litoralis
Amphisbaena barbouri Anolis loysianus
Amphisbaena carlgansi Anolis lucius
Amphisbaena cubana Anolis luteogularis luteogularis
Cadea blanoides Anolis luteogularis calceus
Anolis agueroi Anolis luteogularis jaumei
Anolis ahli Anolis luteogularis nivevultus
Anolis alayoni Anolis macilentus
Anolis alfaroi Anolis mestrei
Anolis allisoni Anolis noblei noblei
Anolis allogus Anolis noblei galeifer
Anolis altitudinalis Anolis ophiolepis
Anolis alutaceus Anolis oporinus
Anolis anfiloquioi Anolis paternus paternus
Anolis angusticeps angusticeps Anolis paternus pinarensis
Anolis argenteolus Anolis porcatus porcatus
Anolis argillaceus Anolis porcatus aracelyae
Anolis baracoae Anolis porcus
Anolis barbatus Anolis pumilus
Anolis bartschi Anolis quadriocellifer
Anolis birama Anolis rejectus
Anolis bremeri bremeri Anolis rubribarbus
Anolis centralis Anolis ruibali
Anolis chamaeleonides Anolis sagrei sagrei
Anolis clivicola Anolis sagrei greyi
Anolis confusus Anolis smallwoodi smallwoodi
Anolis cupeyalensis Anolis smallwoodi palardis
Anolis cyanopleurus cyanopleurus Anolis smallwoodi saxuliceps
Anolis cyanopleurus orientalis Anolis spectrum
Anolis delafuentei Anolis terueli
Anolis equestris equestris Anolis toldo
Anolis equestris buidei Anolis turquinensis
Anolis equestris juraguensis Anolis vanidicus
Anolis equestris persparsus Anolis vermiculatus
Anolis equestris thomasi Anolis vescus
Anolis equestris verreonensisi Aristelliger reyesi
Anolis fugitivus Cricosaura typica
Anolis garridoi Cyclura nubila
Anolis guafe Diploglossus delasagra
Anolis guamuhaya Diploglossus garridoi
116 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Diploglossus nigropunctatus Sphaerodactylus ramsdeni

Gonatodes albogularis fuscus Sphaerodactylus richardi
Gymnophthalmus underwoodi (I) Sphaerodactylus ruibali
Hemidactylus haitianus Sphaerodactylus scaber
Hemidactylus frenatus (I) Sphaerodactylus schwartzi
Hemidactylus mabouia (I?) Sphaerodactylus siboney
Hemidactylus turcicus (I) Sphaerodactylus storeyae
Leiocephalus carinatus carinatus Sphaerodactylus torrei torrei
Leiocephalus carinatus aquarius Sphaerodactylus torrei spielmani
Leiocephalus carinatus labrossytus Tarentola americana americana
Leiocephalus carinatus mogotensis Tarentola crombiei
Leiocephalus carinatus zayasis Arrhyton ainictum
Leiocephalus cubensis cubensis Arrhyton dolichura
Leiocephalus macropus macropus Arrhyton procerum
Leiocephalus macropus aegialus Arrhyton redimitum
Leiocephalus macropus asbolomus Arrhyton supernum
Leiocephalus macropus felinoi Arrhyton taeniatum
Leiocephalus macropus hoplites Arrhyton tanyplectum
Leiocephalus macropus hyacinthurus Arrhyton vittatum
Leiocephalus macropus immaculatus Caraiba andreae andreae
Leiocephalus macropus koopmani Caraiba andreae orientalis
Leiocephalus macropus lenticulatus Caraiba andreae peninsulae
Leiocephalus macropus phylax Cubophis cantherigerus cantherigerus
Leiocephalus macropus torrei Cubophis cantherigerus adspersus
Leiocephalus onaneyi Cubophis cantherigerus pepei
Leiocephalus raviceps raviceps Cubophis cantherigerus schwartzi
Leiocephalus raviceps delavarai Epicrates angulifer
Leiocephalus raviceps jaumei Nerodia clarkii compressicauda
Leiocephalus raviceps klinikowskii Tretanorhinus variabilis variabilis
Leiocephalus raviceps uzzelli Tretanorhinus variabilis binghami
Leiocephalus stictigaster stictigaster Tretanorhinus variabilis wagleri
Leiocephalus stictigaster celeustes Tropidophis celiae
Leiocephalus stictigaster gibarensis Tropidophis feicki
Leiocephalus stictigaster lipomator Tropidophis fuscus
Leiocephalus stictigaster lucianus Tropidophis galacelidus
Leiocephalus stictigaster naranjoi Tropidophis hardyi
Leiocephalus stictigaster ophiplacodes Tropidophis hendersoni
Leiocephalus stictigaster sierrae Tropidophis maculatus
Sphaerodactylus argus Tropidophis melanurus melanurus
Sphaerodactylus armasi Tropidophis melanurus dysodess
Sphaerodactylus bromeliarum Tropidophis morenoi
Sphaerodactylus celicara Tropidophis nigriventris
Sphaerodactylus cricoderus Tropidophis pardalis
Sphaerodactylus dimorphicus Tropidophis pilsbryi
Sphaerodactylus docimus Tropidophis semicinctus
Sphaerodactylus elegans elegans Tropidophis spiritus
Sphaerodactylus intermedius Tropidophis wrighti
Sphaerodactylus nigropunctatus alayoi Tropidophis xantogaster
Sphaerodactylus nigropunctatus granti Typhlops anchaurus
Sphaerodactylus nigropunctatus lissodesmus Typhlops anousius
Sphaerodactylus nigropunctatus strategus Typhlops arator
Sphaerodactylus notatus atactus Typhlops contorhinus
Sphaerodactylus ocujal Typhlops golyathi
Sphaerodactylus oliveri Typhlops lumbricalis
Sphaerodactylus pimienta Typhlops notorachius
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 117

Typhlops oxyrhinus Cayo las Brujas (23.63°/-79.13°) (7.23 km2)

Typhlops pachyrhinus Eleutherodactylus atkinsi atkinsi
Typhlops perimychus Osteopilus septentrionalis
Typhlops satelles Ameiva auberi orlandoi
Typhlops cf. silus Anolis angusticeps angusticeps
Caiman crocodilus (I) Anolis equestris brujensis
Crocodylus acutus Anolis jubar santamariae
Crocodylus rhombifer Anolis porcatus porcatus
Anolis pumilus
KEYS OFF THE NORTHERN COAST OF Anolis sagrei sagrei
CUBA Cyclura nubila
Archipiélago de los Colorados Diploglossus delasagra
Cayo Inés de Soto (22.80°/-83.79) Sphaerodactylus nigropunctatus granti
Anolis angusticeps angusticeps Tarentola americana americana
Anolis porcatus porcatus Cubophis cantherigerus cantherigerus
Anolis sagrei sagrei Epicrates angulifer
Cayo Levisa (22.87°/-83.54) Nerodia clarki compressicauda
Ameiva auberi ssp. Tropidophis melanurus ssp.
Anolis porcatus porcatus Cayo Buba (23.18°/-81.12°)
Anolis sagrei sagrei Ameiva auberi ssp.
Cyclura nubila Anolis sagrei sagrei
Caraiba andreae ssp. Hemidactylus haitianus
Cubophis cantherigerus ssp.
Cayo Caguanes (22.40°/-79.14°)
Cayo la Reina (22.86°/-83.55)
Eleutherodactylus planirostris planirostris
Anolis porcatus porcatus
Eleutherodactylus thomasi
Anolis sagrei sagrei
Osteopilus seprentrionalis
Unnamed Cay Anolis allisoni
Epicrates angulifer Anolis alutaceus
Archipiélago de Sabana Anolis angusticeps angusticeps
Cayo Aguado (22.46°/-79.32) Anolis jubar ssp.
Eleutherodactylus planirostris planirostris Anolis lucius
Anolis sagrei sagrei Anolis porcatus porcatus
Anolis lucius Anolis sagrei sagrei
Cyclura nubila Hemidactylus haitianus
Caraica andreae spp. Sphaerodactylus elegans elegans
Cayo Agustín (22.64°/-79.13) Sphaerodactylus nigropunctatus ssp.
Anolis sagrei sagrei Sphaerodactylus scaber
Cyclura nubila Cubophis cantherigerus ssp.
Crocodylus acutus Tropidophis maculatus
Typhlops lumbricalis
Cayo Bahía de Cádiz (23.22°/-80.47°) (6.37 km2)
Ameiva auberi extraria Cayo Caimán de la Bella (22.58°/-78.85°) (0.28 km2)
Anolis homolechis spp. Anolis sagrei sagrei
Anolis jubar spp. Cayo Caimán Grande (= Caimán del Faro) (22.69°/-78.89°)
Cyclura nubila
(0.23 km2)
Diploglossus delasagra
Ameiva auberi orlandoi
Cubophis cantherigerus cantherigerus
Anolis sagrei sagrei
Cayos Blancos (23.25°/-81.00°) (6.12 km2) Sphaerodactylus nigropunctatus granti
Anolis sagrei sagrei Sphaerodactylus scaber
Cayo Blanquizal (23.15°/-80.41°) Cayos Careneros (22.93°/-79.78°) (0.32 km2)
Cyclura nubila Ameiva auberi extraria
Cayo Boca de Sagua (22.95°/-80.02°) Cyclura nubila
Ameiva auberi spp. Cubophis cantherigerus ssp.
118 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Cayos Cobos (23.63°/-79.13°) (9.09 km2) Cayo Español Adentro (22.62°/-79.13°)

Anolis sagrei sagrei Anolis sagrei sagrei
Cyclura nubila Tarentola americana americana
Cayo Conuco (22.55°/-79.47°) (2.90 km2) Cyclura nubila
Osteopilus septentrionalis Cayo Fábrica (22.41°/-79.19°)
Amphisbaena cubana Anolis jubar ssp.
Anolis homolechis
Cayo Felipe (22.63°/-78.63°) (0.05 km2)
Anolis jubar ssp.
Anolis sagrei sagrei
Anolis porcatus porcatus
Anolis sagrei sagrei Cayo Flamenco (22.03°/-81.58°)
Cyclura nubila Anolis sagrei sagrei
Leiocephalus carinatus ssp. Cayo Fragoso (22.75°/-79.50°) (82.41 km2)
Sphaerodactylus elegans elegans Anolis angusticeps angusticeps
Sphaerodactylus nigropunctatus granti Anolis jubar ssp.
Sphaerodactylus scaber Anolis sagrei sagrei
Cubophis cantherigerus cantherigerus Cyclura nubila
Cayo Cruz del Padre (23.27°/-80.91°) Hemidactylus haitianus
Anolis jubar ssp. Sphaerodactylus elegans elegans
Anolis sagrei sagrei Sphaerodactylus notatus ssp.
Cyclura nubila Tarentola americana americana
Caraiba andreae morenoi
Cayo Cueva (22.42°/-79.18°)
Cubophis cantherigerus ssp.
Anolis jubar ssp.
Crocodylus acutus
Anolis lucius
Cayo Francés (22.64°/-79.22°) (6.22 km2)
Cayo Cupey (in Cárdenas’ Bay) (23.07°/-81.13°) (0.96 km2)
Osteopilus septentrionalis
Leiocephalus carinatus ssp.
Ameiva auberi orlandoi
Cayos de las Cinco Leguas (23.13°/-80.87°) (37.69 km2) Anolis angusticeps angusticeps
Ameiva auberi gemmea Anolis equestris potior
Amphisbaena cubana Anolis jubar santamariae
Anolis equestris cincoleguas Anolis pigmaesquestris
Anolis homolechis homolechis Anolis porcatus porcatus
Anolis sagrei sagrei Anolis sagrei sagrei
Cyclura nubila Cyclura nubila
Leiocephalus carinatus ssp. Hemydactylus haetianus
Leiocephalus cubensis ssp. Leiocephalus stictigaster septentrionalis
Leiocephalus stictigaster ssp. Sphaerodactylus elegans elegans
Nerodia clarki compressicauda Sphaerodactylus nigropunctatus granti
Cayo Diana (23.18°/-81.11°) (0.10 km2) Tarentola americana americana
Anolis sagrei sagrei Cubophis cantherigerus cantherigerus
Cayo Ensenachos (22.64°/-79.00°) (1.45 km2) Cayo Galindo (23.24°/-80.88°)
Eleutherodactylus atkinsi atkinsi Cyclura nubila
Osteopilus septentrionalis Cayo Lanzanillo (22.94°/-79.99°) (1.70 km2)
Ameiva auberi orlandoi Ameiva auberi extraria
Anolis angusticeps angusticeps Anolis angusticeps angusticeps
Anolis jubar santamariae Anolis sagrei sagrei
Anolis porcatus porcatus Cyclura nubila
Anolis sagrei sagrei Leiocephalus raviceps ssp.
Cyclura nubila
Leiocephalus macropus ssp. Cayo Lucas (22.45°/-79.28°)
Leiocephalus stictigaster septentrionalis Eleutherodactylus planirostris planirostris
Sphaerodactylus elegans elegans Eleutherodactylus thomasi
Spherodactylus intermedius Anolis jubar ssp.
Tarentola americana americana Anolis lucius
Cubophis cantherigerus ssp. Anolis sagrei sagrei
Epicrates angulifer Leiocephalus stictigaster ssp.
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 119

Cyclura nubila Leiocephalus macropus ssp.

Tarentola americana americana Leiocephalus stictigaster septentrionalis
Cayo las Tocineras (22.90°/-79.72°) (0.15 km2) Sphaerodactylus elegans elegans
Ameiva auberi extraria Sphaerodactylus nigropunctatus granti
Cyclura nubila Tarentola americana americana
Caraiba andreae morenoi
Cayo Monos de Jutía (including Cayos Monitos de Jutía) Cubophis cantherigerus cantherigerus
(22.96°/-79.86°) (0.27 km2) Epicrates angulifer
Ameiva auberi extraria Tretranorhinus variabilis variabilis
Anolis sagrei sagrei Tropidophis melanurus melanurus
Cyclura nubila
Cayo Tío Pepe (22.95°/-79.82°) (5.75 km2)
Sphaerodactylus nigropunctatus granti
Ameiva auberi extraria
Cayo Obispo (22.40°/-7915°) Anolis sagrei sagrei
Cyclura nubila Cyclura nubila
Cayo Palma (22.39°/-79.10°) (0.27 km2) Cayuelo de la Vela (22.94°/-79.76°) (0.02 km2)
Anolis equestris ssp. Ameiva auberi extorris
Anolis jubar ssp.
Cayo Verde (23.14°/80.23°)
Anolis lucius
Cyclura nubila
Cyclura nubila
Archipiélago de Camagüey
Cayo Piedra del Obispo (23.10°/-80.20°)
Cayo Alto (22.30°/-78.31°) (0.97 km2)
Cyclura nubila
Trachemys decussata decussata
Cayo Punta de Piedra (22.59°/-78.58°) (0.11 km2) Ameiva auberi ssp.
Ameiva auberi extrorris Anolis jubar ssp.
Cyclura nubila Anolis sagrei sagrei
Cayo Salinas (22.42°/-79.24°) (1.08 km2) Anolis porcatus porcatus
Eleutherodactytlus thomasi Sphaerodactylus nigropunctatus ssp.
Anolis sagrei sagrei Cyclura nubila
Anolis angusticeps angusticeps Leiocephalus stictigaster ssp.
Anolis jubar ssp. Cubophis cantherigerus ssp.
Anolis lucius Cayos los Ballenatos (in Nuevitas’ Bay) (22.52°/-77.23°)
Cyclura nubila (0.65 km2 for all three cays)
Tarentola americana americana Amphisbaena cubana
Leiocephalus stictigaster ssp. Anolis allisoni
Amphisbaena cubana Anolis jubar balaenarum
Cayo Santa María (22.66°/-79.00°) (21.93 km2) Anolis porcatus porcatus
Eleutherodactylus auriculatus Anolis sagrei sagrei
Eleutherodactylus planirostris Cyclura nubila
Eleutherodactylus varleyi Gonatodes albogularis fuscus
Osteopilus septentrionalis Sphaerodactylus nigropunctatus granti
Peltophryne peltocephala Sphaerodactylus notatus atactus
Trachemys decussata decussata Cayo Coco (22.49°/-78.50°) (334.52 km2)
Ameiva auberi orlandoi Eleutherodactylus atkinsi atkinsi
Anolis angusticeps angusticeps Eleutherodactylus feichtingeri
Anolis centralis Eleutherodactylus riparius
Anolis equestris potior Eleutherodactylus planirostris
Anolis jubar santamariae Eleutherodactylus thomasi ssp.
Anolis pigmaequestris Osteopilus septentrionalis
Anolis porcatus porcatus Peltophryne empusa
Anolis pumilus Peltophryne gundlachi
Anolis sagrei sagrei Peltophryne peltocephala
Cyclura nubila Trachemys decussata decussata
Diploglossus delasagra Ameiva auberi orlandoi
Gonatodes albogularis fuscus Amphisbaena cubana
Hemidactylus haitianus Anolis angusticeps angusticeps
120 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Anolis allisoni Sphaerodactylus nigropunctatus granti

Anolis pumilus Sphaerodactylus scaber
Anolis equestris cyaneus Tarentola americana americana
Anolis jubar cocoensis Caraiba andreae ssp.
Anolis porcatus porcatus Cubophis cantherigerus ssp.
Anolis sagrei sagrei Cubophis cantherigerus cantherigerus
Cyclura nubila Epicrates angulifer
Diploglossus delasagra Tropidophis pardalis
Hemidactylus haitianus Crocodylus acutus
Leiocephalus carinatus ssp. Cayo Guillermo (22.59°/-78.67°) (15.65 km2)
Leiocephalus cubensis ssp. Ameiva auberi orlandoi
Leiocephalus stictigaster septentrionalis Amphisbaena cubana
Sphaerodactylus elegans elegans Anolis angusticeps angusticeps
Sphaerodactylus nigropunctatus granti
Anolis centralis
Sphaerodactylus notatus atactus
Anolis jubar santamariae
Tarentola americana americana
Anolis sagrei sagrei
Arrhyton taeniatum
Cyclura nubila
Caraiba andreae morenoi
Leiocephalus stictigaster septentrionalis
Cubophis cantherigerus schwartzi
Sphaerodactylus nigropunctatus granti
Epicrates angulifer
Tarentola americana americana
Tetranorhinus variabilis variabilis
Cubophis cantherigerus schwartzi
Tropidophis melanurus ssp.
Typhlops lumbricalis
Tropidophis pardalis
Typhlops satelles (?)
Typhlops lumbricalis
Crocodylus acutus
Crocodylus acutus
Cayo Judas (22.25°/-78.32°)
Cayo Confites (22.19°/-77.66°)
Ameiva auberi ssp.
Anolis sagrei sagrei
Anolis equestris ssp.
Cayo Cruz (22.25°/-77.81°) (26.14 km2) Anolis jubar ssp.
Ameiva auberi orlandoi (?) Anois sagrei sagrei
Anolis jubar ssp. Cyclura nubila
Anolis porcatus porcatus Sphaerodactylus nigropunctatus ssp.
Anolis sagrei sagrei Caraiba andreae ssp.
Cyclura nubila Cubophis cantherigerus ssp.
Leiocephalus stictigaster septentrionalis
Cayo Mégano Grande (22.33°/-77.91°) (7.65 km2)
Cubophis cantherigerus cantherigerus
Ameiva auberi orlandoi
Cayo Guajaba (21.85°/-77.54°) (105.05 km2) Anolis sagrei sagrei
Eleutherodactylus atkinsi atkinsi Leiocephalus stictigaster septentrionalis
Osteopilus septentrionalis
Cayo Paredón de Lado (= Cayo Paredón del Medio)
Peltophryne empusa
(22.48°/-78.21°) (0.02 km2)
Peltophryne gundlachi
Anolis sagrei sagrei
Peltophryne peltocephala
Trachemys decussata decussata Cayo Paredón Grande (22.47°/-78.16°) (10.71 km2)
Ameiva auberi sublesta Eleutherodactylus atkinsi atkinsi
Amphisbaena cubana Osteopilus septentrionalis
Anolis angusticeps angusticeps Peltophryne peltocephala
Anolis centralis Ameiva auberi
Anolis equestris ssp. Amphisbaena cubana
Anolis jubar ssp. Anolis angusticeps angusticeps
Anolis porcatus porcatus Anolis centralis
Anolis sagrei sagrei Anolis homolechis homolechis
Cyclura nubila Anolis jubar ssp.
Hemidactylus haitianus Anolis sagrei sagrei
Leiocephalus stictigaster septentrionalis Cyclura nubila
Sphaerodactylus elegans elegans Leiocephalus stictigaster septentrionalis
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 121

Sphaerodactylus nigropunctatus ssp. Anolis centralis

Sphaerodactylus notatus atactus Anolis equestris sabinalensis
Tarentola americana americana Anolis jubar ssp.
Cubophis cantherigerus schwartzi Anolis porcatus porcatus
Epicrates angulifer Anolis sagrei sagrei
Nerodia clarki compressicauda Cyclura nubila
Tropidophis melanurus Diploglossus delasagra
Tropidophis pardalis Leiocephalus carinatus ssp.
Tropidophis semicinctus Leiocephalus stictigaster ssp.
Typhlops lumbricalis Sphaerodactylus elegans elegans
Cayo Rabihorcado (S of Cayo Coco) (22.40°/-78.43°) Sphaerodactylus notatus atactus
(0.12 km2) Tarentola americana americana
Cyclura nubila Arrhyton taeniatum
Caraiba andreae ssp.
Cayo Romano (22.25°/-78.00°) (680.07 km2) Cubophis cantherigerus ssp.
Eleutherodactylus atkinsi atkinsi Epicrates angulifer
Osteopilus septentrionalis Nerodia clarkii compresicauda
Peltophryne empusa Tretanorhinus variabilis
Peltophryne gundlachi Tropidophis pardalis
Peltophryne peltocephala Typhlops lumbricalis
Trachemys decussata decussata Crocodylus acutus
Ameiva auberi ssp.
Amphisbaena cubana Cayos off the Northern Coast of Holguín Province
Anolis angusticep angusticeps Cayo Saetía (22.75°/-75.50°)
Anolis centralis Eleutherodactylus varleyi
Anolis equestris ssp. Osteopilus septentrionalis
Anolis jubar ssp. Peltophryne peltocephala
Anolis porcatus porcatus Trachemys decussata decussata
Anolis sagrei sagrei Ameiva auberi ssp.
Cyclura nubila Anolis allisoni
Hemidactylus haitianus Anolis equestris ssp.
Leiocephalus carinatus ssp. Anolis homolechis homolechis
Leiocephalus stictigaster ssp. Anolis litoralis
Sphaerodactylus elegans elegans Anolis jubar gibarensis
Sphaerodactylus nigropunctatus granti Anolis porcatus porcatus
Sphaerodactylus notatus atactus Anolis porcus
Caraiba andreae ssp. Anolis sagrei sagrei
Cubohis cantherigerus ssp. Cyclura nubila
Epicrates angulifer Leiocephalius carinatus ssp.
Tropidophis melanurus ssp. Leiocephalus macropus ssp.
Tropidophis pardalis Sphaerodactylus nigropunctatus granti
Tropidophis semicinctus Cubophis cantherigerus schwartzi
Crocodylus acutus Epicrates angulifer

Cayo Sabinal (21.67°/-77.25°) (338.32 km2) KEYS OFF THE SOUTHERN COAST OF
Eleutherodactylus riparius
Eleutherodactylus planirostris
CUBA
Cayos de San Felipe
Osteopilus septentrionalis
Cayo Coco (21.95°/-83.39°)
Peltophryne empusa
Cyclura nubila
Peltophryne gundlachi
Peltophryne peltocephala Cayo Juan García (21.98°/-83.62°)
Trachemys decussata decussata Ameiva auberi sanfelipensis
Ameiva auberi sublesta Anolis angusticeps angusticeps
Amphisbaena cubana Anolis porcatus porcatus
Anolis allisoni Anolis sagrei sagrei
Anolis angusticeps angusticeps Cyclura nubila
122 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Leiocephalus cubensis minor Cayo del Macío (22.15°/-81.61°)

Cubophis cantherigerus cantherigerus Ameiva auberi zugi
Crocodylus acutus Anolis angusticeps angusticeps
Cayo Real (21.96°/-83.62°) Anolis porcatu porcatus
Osteopilus septentrionalis Anolis sagrei sagrei
Ameiva auberi sanfelipensis Cyclura nubila
Anolis angusticeps angusticeps Leiocephalus carinatus ssp.
Anolis homolechis homolechis Leiocephalus cubensis ssp.
Anolis luteogularis sanfelipensis Crocodylus acutus
Anolis porcatus porcatus Cayo Miguel (22.04°/-81.25°)
Anolis sagrei sagrei Ameiva auberi zugi
Cyclura nubila Anolis sagrei sagrei
Dipologlossus delasagra Cyclura nubila
Leiocephalus cubensis minor Leiocephalus carinatus ssp.
Sphaerodactylus notatus atactus
Cayo Pinga Larga (22.06°/-81.28°)
Cubophis cantherigerus cantherigerus
Anolis sagrei sagrei
Tropidophis melanurus
Cyclura nubila
Crocodylus acutus
Leiocephalus carinatus ssp.
Cayo Sijú (21.95°/-83.50°)
Cayo Punta Arena (22.18°/-81.57°)
Ameiva auberi ssp.
Ameiva auberi zugi
Anolis angusticeps angusticeps
Anolis porcatus porcatus Anolis angusticeps angusticeps
Anolis sagrei sagrei Anolis porcatus porcatus
Cyclura nubila Anolis sagei sagrei
Leiocephalus cubensis ssp. Crocodylus acutus
Cubophis cantherigerus ssp. Cayo Sibeque (= Cayo los Pájaros) (22.07°/-81.21°)
Crocodylus acutus Ameiva auberi zugi
Cayos Blancos del Sur Anolis sagrei sagrei
Cayo Caoba (22.12°/-81.65°) Cyclura nubila
Ameiva auberi zugi Leiocephalus cubensis cubensis
Cyclura nubila Cayo del Venado (22.09°/-81.28°)
Leiocephalus cubensis ssp. Ameiva auberi zugi
Cayo Diego Pérez (22.06°/-81.57°) Anolis sagrei sagrei
Ameiva auberi zugi Cyclura nubila
Anolis porcatus porcatus Cubophis cantherigerus ssp.
Anolis sagrei sagrei Archipiélago de los Canarreos
Cayo Ernest Thaelman (22.03°/-81.39°) Cayo Avalos (21.55°/-82.17°)
Ameiva auberi zugi Ameiva auberi zugi
Anolis angusticeps angusticeps Anolis angusticeps angusticeps
Anolis porcatus porcatus Anolis homolechis homolechis
Anolis sagrei sagrei Anolis porcatus porcatus
Cyclura nubila Anolis sagrei sagrei
Leiocephalus carinatus ssp. Cyclura nubila
Leiocephalus cubensis ssp. Sphaerodactylus notatus atactus
Leiocephalus stictigaster ssp. Cubophis cantherigerus cantherigerus
Cubophis cantherigerus ssp. Epicrates angulifer
Epicrates angulifer Crocodylus acutus
Crocodylus acutus Cayos Los Ballenatos (21.57°/-81.67°)
Cayo la Esperanza (22.07°/-81.35°) Anolis sagrei sagrei
Ameiva auberi zugi Cyclura nubila
Anolis sagrei sagrei Sphaerodactylus notatus atactus
Cyclura nubila Cayo Campo (21.57°/-82.30°)
Cayo Longino (22.08°/-81.28°) Ameiva auberi zugi
Anolis sagrei sagrei Anolis angusticeps angusticeps
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 123

Anolis homolechis homolechis Cyclura nubila

Anolis sagrei sagrei Sphaerodactylus oliveri
Cyclura nubila Cubophis cantherigerus cantherigerus
Leiocephalus cubensis pambasileus Epicrates angulifer
Cubophis cantherigerus cantherigerus Tretanorhinus variabilis ssp.
Crocodylus acutus Crocodylus acutus
Cayo Cantiles (21.59°/-82.03°) Cayo Majá (= Cayo los Majáes) (21.53°/-81.60°)
Osteopilus septentrionalis Ameiva auberi zugi
Trachemys decussata ssp. Anolis sagrei sagrei
Ameiva auberi zugi Cyclura nubila
Anolis angusticeps angusticeps Cubophis cantherigerus ssp.
Anolis homolechis homolechis
Cayo Matías (21.57°/-82.43°)
Anolis luteogularis coctilis
Ameiva auberi zugi
Anolis porcatus porcatus
Anolis angusticeps angusticeps
Anolis sagrei sagrei
Anolis porcatus porcatus
Cyclura nubila
Anolis sagrei sagrei
Sphaerodactylus elegans elegans
Cyclura nubila
Sphaerodactylus notatus atactus
Cubophis cantherigerus cantherigerus
Sphaerodactylus storeyae
Crocodylus acutus
Caraiba andreae melopyrrha
Cubophis cantherigerus cantherigerus Cayo Matiítas (21.57°/-82.40°)
Epicrates angulifer Ameiva auberi zugi
Tropidophis melanurus ssp. Anolis angusticeps angusticeps
Crocodylus acutus Anolis porcatus porcatus
Anolis sagrei sagrei
Cayo Estopa (21.65°/-81.80°)
Cyclura nubila
Anolis sagrei sagrei
Cubophis cantherigerus cantherigerus
Cyclura nubila
Crocodylus acutus Crocodylus acutus

Cayo Farito (22.62°/-81.67°) Cayo Pasaje (21.70°/-81.95°)

Anolis sagrei sagrei Anolis angusticeps angusticeps
Cyclura nubila Anolis sagrei sagrei
Crocodylus acutus
Cayo Flamenco (22.02°/-81.58°)
Anolis sagrei sagrei Cayo el Rosario (21.60°/-81.94°)
Ameiva auberi zugi
Cayo Hicacos (21.57°/-82.35°) Anolis angusticeps angusticeps
Ameiva auberi zugi Anolis porcatus porcatus
Anolis angusticeps angusticeps Anolis sagrei sagrei
Anolis porcatus porcatus Cyclura nubila
Anolis sagrei sagrei Leiocephalus carinatus ssp.
Cyclura nubila Cubophis cantherigerus cantherigerus
Leiocephalus cubensis pambasileus Epicrates angulifer
Cubophis cantherigerus cantherigerus Crocodylus acutus
Crocodylus acutus
Isla de la Juventud (= Isla de Pinos) (21.70°/-82.84°)
Cayo Inglés (21.62°/-81.26°) (2,237 km2)
Leiocephalus carinatus ssp. Eleutherodactylus atkinsi atkinsi
Sphaerodactylus oliveri Eleutherodactylus auriculatus
Tarentola americana americana Eleutherodactylus dimidiatus
Cayo Largo del Sur (21.36°/-81.48°) Eleutherodactylus pinarensis
Osteopilus septentrionalis Eleutherodactylus planirostris
Ameiva auberi zugi Eleutherodactylus riparius
Anolis lucius (I?) Eleutherodactylus staurometopon
Anolis luteogularis Eleutherodactylus varleyi
Anolis porcatus porcatus Osteopilus septentrionalis
Anolis sagrei sagrei Lithobates catesbeianus (I)
124 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Peltophryne cataulaciceps Cayos de Ana María

Peltophryne empusa Cayo Arenas (21.48°/-78.81°)
Peltophryne gundlachi Anolis sagrei sagrei
Peltophryne peltocephala Cayo la Cana (21.38°/-78.80°)
Trachemys decussata angusta Anolis sagrei sagrei
Ameiva auberi marcida
Ameiva auberi nigriventris Archipiélago de los Jardines de la Reina
Ameiva auberi secta Cayo Anclitas (20.78°/-78.91°)
Amphisbaena cubana Ameiva auberi galbiceps
Anolis alutaceus Anolis porcatus porcatus
Anolis angusticeps angusticeps Anolis sagrei sagrei
Anolis bremeri insulaepinorum Cyclura nubila
Anolis chamaeleonides Leiocephalus carinatus cayensis
Anolis homolechis homolechis Leiocephalus cubensis paraphrus
Anolis loysianus Sphaerodactylus argus
Anolis luteogularis delacruzi Sphaerodactylus notatus atactus
Anolis luteogularis hassleri Tarentola americana americana
Anolis luteogularis sectilis Cubophis cantherigerus ssp.
Anolis ophiolepis Cayo Boca de Piedra Chica (20.77°/-78.88°)
Anolis paternus paternus Ameiva auberi galbiceps
Anolis porcatus porcatus Leioicephalus carinatus
Anolis pumilus Leiocephalus cubensis paraphrus
Anolis sagrei sagrei
Cayo Boca de Piedra Piloto (= Cayo Piedra Piloto) (20.76°/-
Cadea blanoides
78.83°)
Cadea palirostrata
Ameiva auberi galbiceps
Cyclura nubila
Anolis sagrei sagrei
Diploglossus delasagra
Cyclura nubila
Leiocephalus carinatus microcyon
Leiocephalus carinatus cayensis
Leiocephalus cubensis gigas
Leiocephalus cubensis paraphrus
Leiocephalus macropus ssp.
Leiocephalus stictigaster astictus Cayo Boca Rica (20.61°/-78.48°)
Leiocephalus stictigaster exotheotus Ameiva auberi ssp.
Sphaerodactylos elegans elegans Anolis sagrei sagrei
Sphaerodactylus notatus atactus Cayo Cabahoramas (20.75°/-78.75°)
Sphaerodactylus storeyae Leiocephalus carinatus cayensis
Tarentola americana americana Cubophis cantherigerus ssp.
Arrhyton taeniatum
Arrhyton vittatuum Cayo Caballones (20.88°/-79.01°)
Caraiba andreae nebulatus Ameiva auberi galbiceps
Cubophis cantherigerus cantherigerus Anolis sagrei sagrei
Epicrates angulifer Cyclura nubila
Tretanorhinus variabilis insulaepinorum Leiocephalus carinatus cayensis
Tropidophis maculatus Sphaerodactylus argus
Tropidophis melanurus ericksoni Cayo Cabeza del Este (20.57°/-78.33°)
Tropidophis pardalis Ameiva auberi galbiceps
Typhlops lumbricalis Anolis sagrei sagrei
Caiman crocodilus (I) Cyclura nubila
Crocodylus acutus Sphaerodactylus argus
Crocodylus rhombifer Tarentola americana americana
Keys off the Southern Coast of Sancti Spíritus Province Cayos Cachiboca (including unnamed cay 3 km NW)
Cayo Macho de Tierra (= Cayo Macho de Afuera) (21.59°/- (20.67°/-78.75°)
79.78°) Ameiva auberi galbiceps
Ameiva auberi atrothorax Anolis sagrei sagrei
Anolis sagrei sagrei Cyclura nubila
Cyclura nubila Leiocephalus carinatus cayensis
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 125

Leiocephalus cubensis paraphrus Unnamed Cayo (E of Cayo Juan Grín) (20.63°/-78.53°)

Sphaerodactylus argus Sphaerodactylus argus
Tarentola americana americana Sphaerodactylus elegans elegans
Cubophis cantherigerus ssp. Cayos del Golfo de Guacanayabo
Cayo Caguamas (= Cayo la Tronconera) (20.58°/-78.38°) Cayo Blanco (20.48°/-77.98°)
Anolis sagrei sagrei Anolis sagrei sagrei
Cyclura nubila Cyclura nubila
Sphaerodactylus argus Cayos de Sevilla (20.62°/-77.44°)
Tarentola americana americana Cyclura nubila
Cayo Campos (21.54°/-78.75°) Keys off the Southern Coast of Santiago de Cuba
Anolis sagrei sagrei Province
Cayo Camposanto (20.75°/-78.75°) Cayo Damas (19.97°/-76.40°)
Ameiva auberi galbiceps Anolis jubar oriens
Anolis sagrei sagrei Leiocephalus macropus inmaculatus
Cyclura nubila Sphaerodactylus dimorphicus
Cayo Granada (20.63°/-78.25°)
Leiocephalus carinatus cayensis
Cayo Grande (= Cayo Boca Grande) (20.94°/-79.13°)
Ameiva auberi galbiceps
Anolis sagrei sagrei
Cyclura nubila
Leiocephalus carinatus cayensis
Sphaerodactylus elegans elegans
Cubophis cantherigerus ssp.
Cayo Juan Grín (20.63°/-78.53°)
Ameiva auberi galbiceps
Anolis sagrei sagrei
Leiocephalus carinatus cayensis
Leiocephalus cubensis paraphrus
Sphaerodactylus argus
Cayo las Cruces (22.75°/-78.78°)
Ameiva auberi galbiceps
Anolis sagrei sagrei
Leiocephalus cubensis paraphrus
Cayos Levisa (20.53°/-78.09°)
Sphaerodactylus argus
Tarentola americana americana
Cayo Miraflores (20.75°/-78.75°)
Anolis sagrei sagrei
Cyclura nubila
Cayo Piedra Grande (22.75°/-78.82°)
Ameiva auberi galbiceps
Anolis sagrei sagrei
Cyclura nubila
Cayo Puerto Escondido (20.75°/-78.75°)
Anolis sagrei sagrei
Cayo Rosquete (20.57°/-78.22°)
Anolis jubar ssp.
Tarentola americana americana
126 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

JAMAICA

Byron Wilson1, Thera Edwards2, and Robert Powell

Over 65 species of amphibians and reptiles have latitude in decimal degrees. The attribute table
been documented on Jamaica and its satellites. was exported to Excel and the data converted to
Several species are extinct, many are threatened return results to two decimal places, and change
with extinction, and several others are not native hectares to square kilometres. The calculated areas
to the area, but have become established on islands were verified by selecting two random cays/islands
due to human activity. Besides sources noted in and comparing those results to measures generated
the Introduction, we relied on the overviews of using the online Google Planimeter.
Crombie (1999) and Wilson (2011) and a treatise
by Lazell (1996). JAMAICA BANK
Perimeters of cays and offshore islands were Jamaica (18.15°/-77.30°) (10,829 km2)
derived from tracing land areas, using either Garmin Eleutherodactylus alticola
Eleutherodactylus andrewsi
MapSource or Google Earth®. The preferred source Eleutherodactylus cavernicola
was Google Earth®, as it provided a current satellite Eleutherodactylus cundalli
image of the land mass. Cays and islands that were Eleutherodactylus fuscus
not visible on Google Earth® were digitized in Eleutherodactylus glaucoreius
MapSource using BlueChart Americas v9.5 maps Eleutherodactylus gossei gossei
Eleutherodactylus gossei oligaulax
(Chart Gb383730, Chart Us26125 Portland Bight, Eleutherodactylus grabhami
Chart Us26127 (B) Approaches to Kingston, Morant Eleutherodactylus griphus
Cays). In Map Source, the grid was set to UTM and Eleutherodactylus jamaicensis
the datum to WGS 84. The outline of the cay/island Eleutherodactylus johnstonei (I)
was traced and saved as a dxf file. CAD2Shape Eleutherodactylus junori
Eleutherodactylus luteolus
software was used to convert the dxf file to an Eleutherodactylus nubicola
ESRI ArcMap compatible shapefile. The shapefile Eleutherodactylus orcutti
was opened in a data frame set to UTM Zone Eleutherodactylus pantoni amiantus
18N and assigned a UTM projection in ArcMap. Eleutherodactylus pantoni pantoni
The file was then re-projected into the JAD2001 Eleutherodactylus pentasyringos
Eleutherodactylus planirostris (I)
datum. Cays and islands with perimeters traced in Eleutherodactylus sisyphodemus
Google Earth® were saved as kml files. A JAD2001 Leptodactylus fallax (I, E)
dataframe set at decimal degrees and metric units Lithobates catesbeianus (I)
was opened and the kml files and JAD2001 files Osteopilus crucialis
were added. All individual shapefiles were merged Osteopilus marianae
Osteopilus ocellatus
into one shapefile. The XTools extension was Osteopilus sp.
used to calculate the area of each island/cay in Osteopilus wilderi
hectares in the final merged shapefile. The central Rhinella marina (I)
point of each island/cay was calculated using the Trachemys terrapen
“convert features to spherical centroids” tool in Ameiva dorsalis
Anolis garmani
the Graphics and Shapes extension. Since the data Anolis grahami aquarum
frame was set to decimal degrees the centroid’s X Anolis grahami grahami
and Y coordinates were returned as longitude and Anolis lineatopus ahenobarbus
1
Department of Life Sciences, University of the West Indies, Mona, Jamaica <[email protected]>
2
Department of Geography and Geology, University of the West Indies, Mona, Jamaica <[email protected]>
3
Department of Biology, Avila University, Kansas City, Missouri 64145-1698, USA <[email protected]>
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 127

Anolis lineatopus lineatopus Sphaerodactylus goniorhynchus

Anolis lineatopus merope Sphaerodactylus oxyrhinus
Anolis lineatopus neckeri Sphaerodactylus parkeri
Anolis opalinus Sphaerodactylus richardsoni gossei
Anolis reconditus Sphaerodactylus richardsoni richardsoni
Anolis sagrei ssp. (I?) Sphaerodactylus semasiops
Anolis valencienni Sphaerodactylus sp.
Aristelliger praesignis praesignis Spondylurus fulgidus
Aristelliger titan (F) Tarentola albertschwartzi (PE)
Celestus barbouri Epicrates subflavus
Celestus crusculus crusculus Hypsirhynchus ater (PE)
Celestus crusculus cundalli
Hypsirhynchus callilaemus
Celestus duquesneyi
Hypsirhynchus funererus
Celestus fowleri
Hypsirhynchus polylepis
Celestus hewardii
Celestus microblepharis (PE) Tropidophis jamaicensis
Celestus molesworthi (PE) Tropidophis stejnegeri
Celestus occiduus (PE) Tropidophis stullae
Cyclura collei Typhlops jamaicensis
Gonatodes albogularis notatus (I?) Crocodylus acutus
Hemidactylus mabouia (I?) Big Halfmoon Cay (17.74°/-77.09°) (0.04 km2)
Leiocephalus jamaicensis (F) Ameiva dorsalis
Sphaerodactylus argus
Sphaerodactylus dacnicolor Big Pelican Cay (17.80°/-77.02°) (0.03 km2)
Sphaerodactylus gilvitorques (PE) Ameiva dorsalis

Figure 6. Map of Jamaica and surrounding small islands and cays.

128 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Bogue Islands (18.45°/-77.94°) (0.16 km2 for three cays

combined)
Aristelliger praesignis praesignis
Cabarita Island (18.37°/-76.88°) (0.02 km2)
Anolis grahami grahami
Anolis lineatopus merope
Sphaerodactylus goniorhynchus
Careening Island (= Crane Island) (17.88°/-77.08°) (0.21 km2)
Anolis grahami grahami
Anolis lineatopus lineatopus
Goat Island (= Great Goat Island) (17.87°/-77.05°) (2.20 km2)
Anolis grahami grahami
Anolis lineatopus lineatopus
Anolis opalinus
Cyclura collei (PE)
Sphaerodactylus argus
Epicrates subflavus (?)
Lime Cay (17.92°/-76.82°) (0.02 km2)
Ameiva dorsalis
Little Goat Island (17.89°/-77.07°) (1.04 km2)
Anolis grahami grahami
Anolis lineatopus lineatopus
Anolis opalinus
Cyclura collei (PE)
Sphaerodactylus argus
NE Morant Cay (17.42°/-75.97°) (0.07 km2)
Aristelliger praesignis praesignis
Sphaerodactylus notatus atactus (I?)
NE Pedro Cay (= Top Cay) (17.05°/-77.75°) (0.04 km2)
Sphaerodactylus argus
Pigeon Island (17.80°/-77.07°) (0.18 km2)
Ameiva dorsalis
SE Morant Cay (17.39°/-75.98°) (0.09 km2)
Aristelliger praesignis praesignis
SW Morant Cay (= Rocky Morant Cay) (17.39°/-76.00
(0.02 km2)
Aristelliger praesignis praesignis
SW Pedro Cay (= Bird Cay) (16.99°/-77.81°) (0.17 km2)
Aristelliger praesignis praesignis
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 129

HISPANIOLA AND NAVASSA

Robert Powell1
Well over 200 species of amphibians and reptiles NAVASSA BANK
have been documented on Hispaniola and its Navassa Island (18.40°/-75.01°) (5.2 km2)
satellites, and an additional nine species occur or Chelonoidis sp. (F)
Anolis longiceps
are thought to have occurred on Navassa. Several Aristelliger cochranae
species are extinct, many are threatened with Celestus badius
extinction, and several others are not native to the Cyclura onchiopsis (PE)
area, but have become established on islands due to Leiocephalus eremitus (PE)
human activity. Sphaerodactylus becki
Tropidophis bucculentus (PE)
Besides sources noted in the Introduction, I Typhlops sulcatus (PX)
relied on the overviews of Powell et al. (1999) and
Powell and Incháustegui (2011), various reports HISPANIOLA BANK
of my own fieldwork (including a trip to Navassa; Hispaniola (18.95°/-71.26°) (73,929 km2)
Eleutherodactylus abbotti
Powell 1999), and personal communications
Eleutherodactylus alcoae
with Sixto J. Incháustegui. I gleaned latitude and Eleutherodactylus amadeus
longitude from the online gazetteer (www.tageo. Eleutherodactylus aporostegus
com) and gazetteers prepared by the Defense Eleutherodactylus apostates
Mapping Agency Topographic Center (1973). Eleutherodactylus armstrongi
Eleutherodactylus audanti audanti
For areas of the Cayos Siete Hermanos and Cayo
Eleutherodactylus auriculatoides
Pisaje, I consulted Burns et al. (1992) and Yeska et Eleutherodactylus bakeri
al. (2000), respectively. Eleutherodactylus bothroboans

Figure 7. Map of the Hispaniola and Navassa banks.

1
Department of Biology, Avila University, Kansas City, Missouri 64145-1698, USA <[email protected]>
130 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Eleutherodactylus brevirostris Eleutherodactylus warreni

Eleutherodactylus caribe Eleutherodactylus weinlandi weinlandi
Eleutherodactylus chlorophenax Eleutherodactylus weinlandi chersonesodes
Eleutherodactylus corona Eleutherodactylus wetmorei wetmorei
Eleutherodactylus coqui (W?) Eleutherodactylus wetmorei ceraemerus
Eleutherodactylus counouspeus Hypsiboas heilprini
Eleutherodactylus darlingtoni Leptodactylus albilabris
Eleutherodactylus diplasius Osteopilus dominicensis
Eleutherodactylus dolomedes Osteopilus pulchrilineatus
Eleutherodactylus eunaster Osteopilus vastus
Eleutherodactylus flavescens Lithobates catesbeianus (I)
Eleutherodactylus fowleri Peltophryne fluviatica (PE)
Eleutherodactylus furcyensis Peltophryne fracta
Eleutherodactylus glandulifer Peltophryne guentheri
Eleutherodactylus glanduliferoides Rhinella marina (I)
Eleutherodactylus glaphycompus Chelonoidis sp. (F)
Eleutherodactylus grahami Trachemys decorata
Eleutherodactylus haitianus Trachemys scripta elegans (I)
Eleutherodactylus heminota Trachemys stejnegeri vicina
Eleutherodactylus hypostenor Ameiva chrysolaema chrysolaema
Eleutherodactylus inoptatus Ameiva chrysolaema alacris
Eleutherodactylus jugans Ameiva chrysolaema boekeri
Eleutherodactylus lamprotes Ameiva chrysolaema defensor
Eleutherodactylus leoncei Ameiva chrysolaema evulsa
Eleutherodactylus limbensis Ameiva chrysolaema ficta
Eleutherodactylus lucioi Ameiva chrysolaema jacta
Eleutherodactylus melatrigonum Ameiva chrysolaema parvoris
Eleutherodactylus minutus Ameiva chrysolaema procax
Eleutherodactylus montanus Ameiva chrysolaema quadrijugis
Eleutherodactylus nortoni Ameiva chrysolaema regularis
Eleutherodactylus notidodes Ameiva chrysolaema umbratilis
Eleutherodactylus oxyrhyncus Ameiva lineolata lineolata
Eleutherodactylus parabates Ameiva lineolata meracula
Eleutherodactylus paralius Ameiva lineolata perplicata
Eleutherodactylus parapelates Ameiva lineolata privigna
Eleutherodactylus patriciae Ameiva taeniura azuae
Eleutherodactylus paulsoni Ameiva taeniura barbouri
Eleutherodactylus pictissimus pictissimus Ameiva taeniura ignobilis
Eleutherodactylus pictissimus apantheatus Ameiva taeniura meyerabichi
Eleutherodactylus pictissimus eremus Ameiva taeniura regnatrix
Eleutherodactylus pituinus Ameiva taeniura taeniura
Eleutherodactylus poolei Ameiva taeniura tofacea
Eleutherodactylus probolaeus Ameiva taeniura vafra
Eleutherodactylus rhodesi Ameiva taeniura varica
Eleutherodactylus rucillensis Ameiva taeniura vulcanalis
Eleutherodactylus rufifemoralis Anolis aliniger
Eleutherodactylus ruthae Anolis altavelensis
Eleutherodactylus schmidti Anolis alumina
Eleutherodactylus sciagraphus Anolis armouri
Eleutherodactylus semipalmatus Anolis bahorucoensis bahorucoensis
Eleutherodactylus sommeri Anolis bahorucoensis southerlandi
Eleutherodactylus sp. (F) Anolis baleatus altager
Eleutherodactylus thorectes Anolis baleatus baleatus
Eleutherodactylus tychathrous Anolis baleatus caeruleolatus
Eleutherodactylus ventrilineatus Anolis baleatus fraudator
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 131

Anolis baleatus litorisilva Anolis olssoni olssoni

Anolis baleatus multistruppus Anolis olssoni palloris
Anolis baleatus samanae Anolis placidus
Anolis baleatus scelestus Anolis porcatus porcatus (I)
Anolis baleatus sublimis Anolis ricordii leberi
Anolis barahonae albocellatus Anolis ricordii ricordii
Anolis barahonae barahonae Anolis ricordii subsolanus
Anolis barahonae ininquinatus Anolis ricordii viculus
Anolis barahonae mulitus Anolis rimarum
Anolis barbouri Anolis rupinae
Anolis breslini Anolis semilineatus
Anolis brevirostris brevirostris Anolis sheplani
Anolis brevirostris deserticola Anolis shrevei
Anolis brevirostris wetmorei Anolis singularis
Anolis caudalis Anolis strahmi abditus
Anolis chlorocyanus chlorocyanus Anolis strahmi strahmi
Anolis chlorocyanus cyanostictus Anolis websteri
Anolis christophei Anolis whitemani whitemani
Anolis coelestinus coelestinus Anolis whitemani lapidosus
Anolis cristatellus cristatellus (I) Aristelliger expectatus
Anolis cybotes cybotes Aristelliger lar
Anolis cybotes doris Celestus agasepsoides
Anolis cybotes ravifaux Celestus anelpistus (PE)
Anolis darlingtoni Celestus costatus chalcorhabdus
Anolis distichus aurifer Celestus costatus costatus
Anolis distichus dominicensis Celestus costatus leionotus
Anolis distichus favillarum Celestus costatus melanchrous
Anolis distichus ignigularis Celestus costatus neiba
Anolis distichus properus Celestus costatus oreistes
Anolis distichus ravitergum Celestus costatus psychonothes
Anolis distichus suppar Celestus curtissi aporus
Anolis distichus vinosus Celestus curtissi curtissi
Anolis dolichocephalus dolichocephalus Celestus curtissi diastatus
Anolis dolichocephalus portusalus Celestus curtissi hylonomus
Anolis dolichocephalus sarmenticola Celestus darlingtoni
Anolis dominicanus (F) Celestus haetianus haetianus
Anolis etheridgei Celestus haetianus mylicus
Anolis eugenegrahami Celestus haetianus surdus
Anolis fowleri Celestus macrotus
Anolis haetianus Celestus marcanoi
Anolis hendersoni hendersoni Celestus sepsoides
Anolis hendersoni ravidormitans Celestus stenurus alloeides
Anolis insolitus Celestus stenurus stenurus
Anolis koopmani Celestus stenurus rugosus
Anolis longitibialis longitibialis Celestus stenurus weinlandi
Anolis longitibialis specuum Celestus warreni
Anolis marcanoi Cyclura cornuta
Anolis marron Cyclura ricordii
Anolis monticola monticola Gonatodes albogularis notatus (I?)
Anolis monticola quadrisartus Gymnophthalmus underwoodi (W?)
Anolis olssoni alienus Hemidactylus haitianus
Anolis olssoni dominigensis Hemidactylus frenatus (I)
Anolis olssoni extentus Hemidactylus mabouia (I?)
Anolis olssoni ferrugicauda Iguana iguana (W?)
Anolis olssoni montivagus Leiocephalus anonymous (F)
132 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Leiocephalus altavelensis Sphaerodactylus darlingtoni darlingtoni

Leiocephalus apertosculcus (F) Sphaerodactylus darlingtoni mekistus
Leiocephalus barahonensis aureus Sphaerodactylus darlingtoni noblei
Leiocephalus barahonensis barahonensis Sphaerodactylus difficilis difficilis
Leiocephalus barahonensis oxygaster Sphaerodactylus difficilis anthracomus
Leiocephalus endomychus Sphaerodactylus difficilis diolenius
Leiocephalus lunatus arenicolor Sphaerodactylus difficilis lycauges
Leiocephalus lunatus lewisi Sphaerodactylus difficilis peratus
Leiocephalus lunatus lunatus Sphaerodactylus difficilis typhlopous
Leiocephalus lunatus thomasi Sphaerodactylus dommeli (F)
Leiocephalus melanochlorus hypsistus Sphaerodactylus elasmorhynchus
Leiocephalus melanochlorus melanochlorus Sphaerodactylus elegans punctatissimus
Leiocephalus personatus personatus Sphaerodactylus epiurus
Leiocephalus personatus actites Sphaerodactylus ladae
Leiocephalus personatus agraulus Sphaerodactylus lazelli
Leiocephalus personatus budeni Sphaerodactylus leucaster
Leiocephalus personatus elattoprosopon Sphaerodactylus nycteropus
Leiocephalus personatus mentalis Sphaerodactylus ocoae
Leiocephalus personatus poikilometes Sphaerodactylus omoglaux
Leiocephalus personatus pyrrholaemus Sphaerodactylus perissodactylius
Leiocephalus personatus scalaris Sphaerodactylus plummeri
Leiocephalus personatus socoensis Sphaerodactylus randi methorius
Leiocephalus personatus tarachodes Sphaerodactylus randi randi
Leiocephalus personatus trujilloensis Sphaerodactylus randi strahmi
Leiocephalus pratensis chimarus Sphaerodactylus rhabdotus
Leiocephalus pratensis pratensis Sphaerodactylus samanensis
Leiocephalus rhutidira Sphaerodactylus savagei juanilloensis
Leiocephalus schreibersii schreibersii Sphaerodactylus savagei savagei
Leiocephalus semilineatus Sphaerodactylus schuberti
Mabuya hispaniolae (PE) Sphaerodactylus shrevei
Phyllodactylus hispaniolae Sphaerodactylus sommeri
Phyllodactylus sommeri Sphaerodactylus streptophorus sphenophanes
Sphaerodactylus altavelensis altavelensis (?) Sphaerodactylus streptophorus streptophorus
Sphaerodactylus altavelensis brevirostratus Sphaerodactylus thompsoni
Sphaerodactylus altavelensis enriquilloensis Sphaerodactylus williamsi
Sphaerodactylus altavelensis lucioi Sphaerodactylus zygaena
Sphaerodactylus ariasae Spondylurus haitiae (PE)
Sphaerodactylus armstrongi armstrongi Spondylurus lineolatus (PE)
Sphaerodactylus armstrongi hypsinephes Amphisbaena caudalis
Sphaerodactylus asterulus Amphisbaena hyporissor hyporissor
Sphaerodactylus callocricus Amphisbaena hyporissor leberi
Sphaerodactylus cinereus cinereus Amphisbaena innocens
Sphaerodactylus cinereus stejnegeri Amphisbaena leali
Sphaerodactylus clenchi apocoptus Amphisbaena manni
Sphaerodactylus clenchi clenchi Boa constrictor (W)
Sphaerodactylus cochranae Epicrates fordii agametus
Sphaerodactylus copei astreptus Epicrates fordii fordii
Sphaerodactylus copei cataplexis Epicrates gracilis gracilis
Sphaerodactylus copei copei Epicrates gracilis hapalus
Sphaerodactylus copei enochrus Epicrates striatus exagistus
Sphaerodactylus copei pelates Epicrates striatus striatus
Sphaerodactylus copei picturatus Haitiophis anomalus
Sphaerodactylus copei websteri Hypsirhynchus ferox ferox
Sphaerodactylus cryphius Hypsirhynchus melanichnus (PE)
Sphaerodactylus darlingtoni bobilini Hypsirhynchus parvifrons lincolni
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 133

Hypsirhynchus parvifrons niger Cayo Monte Grande (Siete Hermanos) (19.88°/-71.79°)

Hypsirhynchus parvifrons paraniger (0.12 km2)
Hypsirhynchus parvifrons parvifrons Ameiva chrysolaema regularis
Hypsirhynchus parvifrons protenus Anolis cybotes cybotes
Hypsirhynchus scalaris Aristelliger lar
Ialtris agyrtes Celestus stenurus rugosus
Ialtris dorsalis Sphaerodactylus difficilis typhlopous
Ialtris haetianus haetianus Cayo Muertos (Siete Hermanos) (19.85°/-71.82°) (0.12 km2)
Ialtris haetianus perfector Ameiva chrysolaema regularis
Ialtris haetianus vaticinata Leiocephalus schreibersii schreibersii
Ialtris parishi Sphaerodactylus difficilis typhlopous
Mitophis asbolepis
Mitophis calypso Cayo Pascual (Bahía de Samaná) (= Cayo la Farola = Isla
Mitophis leptepileptus Pascal) (19.17°/-69.27)
Mitophis pyrites Anolis baleatus samanae
Tropidophis haetianus haetianus Sphaerodactylus difficilis diolenius
Tropidophis haetianus hemerus Cayo Pisaje (17.84°/-71.28°) (0.29 km2)
Tropidophis haetianus tiburonensis Anolis brevirostris wetmorei
Typhlops agoralionis Aristelliger expectatus
Typhlops capitulatus Leiocephalus barahonensis oxygaster
Typhlops eperopeus Sphaerodactylus randi strahmi
Typhlops gonavensis Cayo Ratas (Siete Hermanos) (19.88°/-71.82°) (0.13 km2)
Typhlops hectus Leiocephalus schreibersii schreibersii
Typhlops proancylops
Typhlops pusillus Cayo Tercero (Siete Hermanos) (19.89°/-71.81°) (0.15 km2)
Typhlops schwartzi Ameiva chrysolaema regularis
Typhlops sulcatus Anolis distichus dominicensis
Typhlops syleptor Leiocephalus schreibersii schreibersii
Typhlops syntherus Cayo Tororú (Siete Hermanos) (19.83°/-71.80°) (0.12 km2)
Typhlops tetrathyreus Ameiva chrysolaema regularis
Typhlops titanops Celestus costatus ssp.
Uromacer catesbyi catesbyi Leiocephalus schreibersii schreibersii
Uromacer catesbyi hariolatus Grosse Caye (18.22°/-73.40°)
Uromacer catesbyi pampineus Ameiva chrysolaema evulsa
Uromacer frenatus frenatus Ameiva taeniura taeniura
Uromacer frenatus chlorauges Leiocephalus personatus elattoprosopon
Uromacer oxyrhynchus Hypsirhynchus parvifrons parvifrons
Crocodylus acutus Uromacer frenatus frenatus
Cayo Arenas (Siete Hermanos) (19.87°/-71.85°) (0.04 km2) Île à Cabrit (Baie de Port-au-Prince) (= Caye Carenage)
None (18.70°/-72.38°)
Cayo Hondo (Bahía de Samaná) (= Cayo Alcatraz = Cayo Ameiva chrysolaema chrysolaema
del Sur = Cayo Playa Hondo) (19.10°/-69.16°) Ameiva lineolata lineolata
Anolis baleatus samanae Anolis brevirostris brevirostris
Anolis caudalis
Cayo Levantado (Bahía de Samaná) (19.10°/-69.17°) (1.5 km2)
Anolis cybotes cybotes
Aristelliger lar
Anolis olssoni extentus
Cayo Monte Chico (Siete Hermanos) (19.86°/-71.77°) Anolis semilineatus
(0.09 km2) Aristelliger expectatus
Ameiva chrysolaema regularis Celestus stenurus weinlandi
Anolis distichus dominicensis Gonatodes albogularis notatus
Aristelliger lar Leiocephalus pratensis chimarus
Leiocephalus personatus scalaris Sphaerodactylus altavelensis brevirostratus
Sphaerodactylus difficilis typhlopous Epicrates fordii manototus
134 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Île de la Gonâve (18.85°/-73.06°) (692 km2) Anolis distichus patruelis

Osteopilus dominicensis Cyclura cornuta (PE)
Ameiva chrysolaema secessa Uromacer catesbyi cereolineatus
Ameiva taeniura barbouri Île de la Petite Gonâve (18.70°/-72.80°)
Anolis caudalis
Anolis caudalis
Anolis chlorocyanus chlorocyanus
Cyclura cornuta
Anolis cybotes doris
Amphisbaena gonavensis
Anolis olssoni insulanus
Hypsirhynchus parvifrons alleni
Anolis singularis
Anolis strahmi abditus Île de la Tortue (20.05°/-72.79°) (188 km2)
Anolis strahmi strahmi Eleutherodactylus inoptatus
Aristelliger expectatus Eleutherodactylus warreni
Celestus curtissi curtissi Osteopilus dominicensis
Celestus sepsoides Ameiva chrysolaema woodi
Gonatodes albogularis notatus Anolis chlorocyanus chlorocyanus
Leiocephalus vinculum Anolis cybotes cybotes
Sphaerodactylus copei deuterus Anolis distichus dominicensis
Sphaerodactylus elegans punctatissimus Anolis semilineatus
Amphisbaena gonavensis Aristelliger expectatus
Epicrates fordii fordii Celestus costatus emys
Epicrates striatus striatus Celestus curtissi diastatus
Hypsirhynchus ferox paracrousis Celestus warreni
Hypsirhynchus parvifrons alleni Cyclura cornuta
Ialtris dorsalis Leiocephalus schreibersii nesomorus
Tropidophis haetianus haetianus Sphaerodactylus difficilis euopter
Typhlops gonavensis Amphisbaena manni
Typhlops pusillus Epicrates striatus warreni
Typhlops sulcatus Haitiophis anomalus
Uromacer catesbyi frondicolor Hypsirhynchus parvifrons tortuganus
Uromacer frenatus dorsalis Ialtris dorsalis
Crocodylus acutus (PX) Ialtris parishi
Tropidophis haetianus haetianus
Île Grande Cayemite (18.61°/-73.75°)
Typhlops pusillus
Osteopilus dominicensis
Uromacer oxyrhynchus
Ameiva taeniura pentamerinthus
Anolis coelestinus demissus Île-à-Vache (18.08°/-73.64°) (48.5 km2)
Anolis cybotes cybotes Eleutherodactylus pictissimus pictissimus
Anolis distichus patruelis Osteopilus dominicensis
Anolis semilineatus Ameiva taeniura aequorea
Celestus sepsoides Anolis coelestinus pecuarius
Celestus stenurus ssp. Anolis cybotes cybotes
Cyclura cornuta Anolis distichus juliae
Sphaerodactylus copei polyommatus Celestus costatus nesobous
Sphaerodactylus elegans punctatissimus Celestus stenurus stenurus
Amphisbaena caudalis Leiocephalus melanochlorus melanochlorus
Amphisbaena cayemite Sphaerodactylus copei cataplexis
Amphisbaena innocens Epicrates striatus exagistus
Hypsirhynchus parvifrons parvifrons Hypsirhynchus parvifrons rosamondae
Typhlops hectus Ialtris dorsalis
Typhlops pusillus Uromacer catesbyi insulaevaccarum
Typhlops sulcatus Uromacer frenatus ssp.
Uromacer catesbyi cereolineatus Crocodylus acutus (PE)
Uromacer frenatus ssp. Isla Alto Velo (17.48°/-71.64°) (1.4 km2)
Île Petite Cayemite (18.61°/-73.81°) Anolis altavelensis
Ameiva taeniura taeniura Aristelliger expectatus
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 135

Leiocephalus altavelensis Leiocephalus lunatus louisae

Sphaerodactylus altavelensis altavelensis Sphaerodactylus savagei juanilloensis
Typhlops sulcatus Epicrates fordii fordii
Isla Beata (17.57°/-71.52°) (42.1 km2) Epicrates striatus striatus
Eleutherodactylus alcoae Hypsirhynchus ferox exedrus
Ameiva chrysolaema abbotti Hypsirhynchus parvifrons stygius
Ameiva lineolata beatensis Typhlops pusillus
Anolis brevirostris wetmorei Uromacer catesbyi inchausteguii
Anolis longitibialis longitibialis Uromacer oxyrhynchus
Aristelliger expectatus
Cyclura cornuta
Leiocephalus barahonensis beatanus
Sphaerodactylus ariasae
Sphaerodactylus thompsoni
Amphisbaena hyporissor leberi
Haitiophis anomalus
Hypsirhynchus parvifrons lincolni
Uromacer frenatus wetmorei
Isla Cabras (Monte Cristi) (19.89°/-71.67°) (0.29 km2)
Ameiva chrysolaema regularis
Ameiva lineolata meracula
Anolis distichus dominicensis
Anolis olssoni olssoni
Anolis whitemani ssp.
Leiocephalus personatus scalaris
Isla Catalina (18.36°/-69.01°) (9.2 km2)
Ameiva chrysolaema parvoris
Ameiva lineolata semota
Anolis cybotes cybotes
Anolis distichus tostus
Anolis semilineatus
Aristelliger lar
Celestus curtissi ssp.
Leiocephalus lunatus melaenascelis
Epicrates fordii fordii
Hypsirhynchus parvifrons paraniger
Typhlops pusillus
Uromacer oxyrhynchus
Isla Catalinita (18.19°/-68.64°) (0.22 km2)
Anolis cybotes ssp.
Celestus costatus aenetergum
Sphaerodactylus savagei ssp.
Isla Saona (18.15°/-68.67°) (105 km2)
Osteopilus dominicensis
Ameiva chrysolaema richardthomasi
Ameiva taeniura rosamondae
Anolis baleatus lineatacervix
Anolis chlorocyanus chlorocyanus
Anolis cybotes ravifaux
Anolis distichus sejunctus
Celestus costatus saonae
Celestus curtissi ssp.
Cyclura cornuta
Hemidactylus haitianus
136 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

PUERTO RICO AND THE VIRGIN ISLANDS

Gregory C. Mayer1

“The multitude of other islands in the group or in part, from that of Puerto Rico, to which they
will well repay the visit of the zoologist, since belong politically. The St. Croix Bank, 40 km south
the life of this section of the West Indies cannot of St. Thomas (on the Puerto Rico Bank), consists
be said to be known until all the cays and islets of St. Croix and four small islets. Its fauna shows
have been investigated” (Wetmore 1927). considerable endemism, but also shares several
Puerto Rico and the Virgin Islands comprise over forms with the Puerto Rico Bank.
150 islands for which herpetological records exist. About 100 species of reptiles and amphibians
These islands are spread over five island banks: have been recorded from the area covered, of which
the Mona Bank, the Monito Bank, the Desecheo more than 20 have been introduced. The introduced
Bank, the Puerto Rico Bank (also known as Greater populations of Cyclura pinguis are deliberate
Puerto Rico: Schmidt 1928), and the St. Croix Bank translocations within the probable former range of
(Fig. 8). The Puerto Rico Bank extends from the this endangered species (Lazell 2005).
Mona Passage in the west to the Anegada Passage The following lists were compiled starting
in the east. On the north, it is bordered by the with the distributional checklist in Mayer (1989),
Puerto Rican Trench, and to the south deep water to which I added the results of more recent
separates it from St. Croix. The islands on this bank fieldwork, data from the examination of museum
are Puerto Rico and adjacent islets, the Passage specimens, and literature records. In addition to
Islands (Vieques and Culebra and associated islets), the general works cited in the introduction, the
the U.S. Virgin Islands (except St. Croix), and the works of Stejneger (1904), Schmidt (1928), Grant
British Virgin Islands. All of these islands were (e.g., 1937), Philibosian and Yntema (1976, 1978),
united into a single land mass during the low sea Rivero (1978, 1998), Heatwole et al. (1981),
level stands associated with the Pleistocene glacial MacLean (1982), and Lazell (1983) are fundamental
maxima, when sea level stood approximately 120 for Puerto Rican/Virgin Island herpetology. More
m below its present level (Heatwole & MacKenzie recent works that have made useful compilations
1967, Fairbanks 1989). Because banks set the limits are Ovaska et al. (2000), Platenberg (2007), Perry
to recent over-land dispersal, they are key entities and Gerber (2011), Platenberg and Boulon (2011),
in herpetogeographic analyses (Barbour & Shreve and Powell et al. (2011). Distributions within the
1935). The islands of the other banks are separated large island of Puerto Rico are addressed in Rivero
from the Puerto Rico Bank by water sufficiently (1978, 1998), Schwartz and Henderson (1991), and
deep to have precluded their being joined to Hedges (2012).
Greater Puerto Rico by any glacial sea level drop. For areas and locations, the chief sources
The Mona Bank, 60 km west of Puerto Rico, has were McGuire (1925), Lazell (1983), the U.S. Geo-
a single island, Isla Mona. Two kilometers to the logical Survey Geographic Names Information
northwest, separated by water over 220 m deep, System (www.geonames.usgs.gov), the Montana
is the Monito Bank, with its single island, Isla State University Environmental Statistics Group
Monito. The Desecheo Bank, 20 km west of Puerto (www.esg.montana.edu), and the National Geo-
Rico, also has but one island, Isla Desecheo. The spatial-Intelligence Agency (http://earth-info.nga.
fauna of these three islands is derived, in whole mil/gns/html/index.html), supplemented by Google

1
Department of Biological Sciences, University of Wisconsin-Parkside, Kenosha, Wisconsin 53141 <[email protected]>
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 137

Figure 8. Map of greater Puerto Rico and adjacent island banks, including the Virgin Islands.

Earth® (www.earth.google.com) and Berkeley- Chelonoidis monensis (F)

Mapper (berkeleymapper.berkeley.edu). Coordin- Ameiva alboguttata
Anolis monensis
ates for the BVI are less precise, as the NGA data Cyclura stejnegeri
are rounded to minutes. Many variant names exist Hemidactylus mabouia (I?)
for many of the islands, and in deciding which Sphaerodactylus monensis
names to use, I have considered official names, Spondylurus monae
navigational charts, local usage, and usage in Borikenophis variegatus
Epicrates monensis
herpetological and other scientific literature. I have Typhlops monensis
not followed any one of these exclusively.
MONITO BANK
ACKNOWLEDGEMENTS Isla Monito (18.16°/-67.95°) (0.16 km2)
Anolis monensis
My primary debt is to James Lazell, who Sphaerodactylus micropithecus
Spondylurus monitae
accompanied me through much of the field work,
and provided support via The Conservation Agency. DESECHEO BANK
The Jarecki family, E.M. Azevedo, D. Ford, R.Z. Isla Desecheo (18.39°/-67.48°) (1.46 km2)
Jenkins, W.P. and E. MacLean, N.R. Mayer, R. Ameiva exsul
Anolis desechensis
Thomas, and J. Walsh assisted in numerous ways,
Sphaerodactylus levinsi
as did K. Tighe and J. Jacobs (National Museum Spondylurus nitidus
of Natural History) and J. Rosado (Museum of Borikenophis portoricensis ssp.
Comparative Zoology) during the most recent
museum work. James Lazell, Gad Perry, Kristiina PUERTO RICO BANK
Puerto Rico (18.25°/-66.50°) (8,674 km2)
Ovaska, and Renata J. Platenberg provided recent Eleutherodactylus antillensis
records. Eleutherodactylus brittoni
Eleutherodactylus cochranae
MONA BANK Eleutherodactylus gryllus
Isla Mona (18.08°/-67.89°) (55.81 km2) Eleutherodactylus hedricki
Eleutherodactylus monensis Eleutherodactylus jasperi
Rhinella marina (I) Eleutherodactylus juanriveroi
138 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Eleutherodactylus karlschmidti Sphaerodactylus townsendi

Eleutherodactylus locustus Spondylurus nitidus
Eleutherodactylus portoricensis Boa constrictor ssp. (I)
Eleutherodactylus richmondi Borikenophis portoricensis portoricensis
Eleutherodactylus unicolor Borikenophis portoricensis prymnus
Eleutherodactylus wightmanae Epicrates inornatus
Hyla cinerea (I) Epicrates granti
Leptodactylus albilabris Eunectes notaeus (W)
Leptodactylus fallax (W) Magliophis stahli
Lithobates grylio (I) Magliophis exiguus subspadix
Lithobates catesbeianus (I) Python bivittatus (W)
Osteopilus septentrionalis (I) Python regius (W)
Peltophryne lemur Python reticulatus (W)
Rhinella marina (I) Python sebae (W)
Scinax ruber (I) Typhlops granti
Trachemys scripta elegans (I) Typhlops hypomethes
Trachemys stejnegeri stejnegeri Typhlops platycephalus
Ameiva exsul Typhlops rostellatus
Ameiva wetmorei Caiman crocodilus (I)
Amphisbaena bakeri
Amphisbaena caeca Cays off the Northern Coast of Puerto Rico
Amphisbaena schmidti Isla de Cabras (= Leper Island) (18.47°/-66.14°)
Amphisbaena xera Ameiva exsul
Anolis cooki Anolis cristatellus cristatellus
Anolis cristatellus cristatellus Anolis pulchellus
Anolis cristatellus cristatellus x A. c. wileyae Punta Salinas Island (18.47°/-66.18°)
Anolis cuvieri Anolis cristatellus cristatellus
Anolis evermanni Anolis pulchellus
Anolis gundlachi
Anolis krugi Cays off the Southern Coast of Puerto Rico
Anolis occultus Caja de Muertos (17.89°/-66.52°)
Anolis poncensis Ameiva exsul
Anolis pulchellus Ameiva wetmorei
Anolis stratulus Amphisbaena xera
Cyclura nubila (I) Anolis cooki
Cyclura pinguis (F*) Anolis cristatellus cristatellus
Diploglossus pleii Anolis pulchellus
Hemidactylus haitianus Phyllodactylus wirshingi
Hemidactylus mabouia (I?) Sphaerodactylus roosevelti
Hemidactylus turcicus (W) Sphaerodactylus townsendi
Iguana delicatissima (W) Borikenophis portoricensis prymnus
Iguana iguana (I) Typhlops granti
Leiocephalus etheridgei (F) Typhlops sp.
Leiocephalus partitus (F) Cayos de Barca (= Cayos de la Barca) (17.92°/-66.24°)
Phyllodactylus wirshingi Anolis cristatellus cristatellus
Scincella lateralis (I)
Sphaerodactylus gaigeae Cayo Cardona (17.96°/-66.63°)
Sphaerodactylus klauberi Ameiva exsul
Sphaerodactylus macrolepis ateles Anolis cristatellus cristatellus
Sphaerodactylus macrolepis grandisquamis Cayo Don Luis (17.94°/-66.97°)
Sphaerodactylus macrolepis guarionex Anolis cooki
Sphaerodactylus macrolepis mimetes
Cayo Punta Aguila (17.95°/-67.21°)
Sphaerodactylus macrolepis phoberus
Anolis cooki
Sphaerodactylus macrolepis spanius
Sphaerodactylus nicholsi Cayos de Ratones (near Jobos) (17.93°/-66.29°)
Sphaerodactylus roosevelti Anolis cristatellus cristatellus
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 139

Cayo Turramote (17.94°/-67.02°) Cayo Palominitos (18.34°/-65.57°)

Sphaerodactylus nicholsi (W) Eleutherodactylus cochranae (W)
Isla Cueva (17.96°/-67.08°) Ameiva exsul
Hemidactylus mabouia (I?) Anolis cristatellus wileyae
Anolis pulchellus (I, E)
Isla Magueyes (17.97°/-67.04°) Anolis stratulus (W)
Rhinella marina (I) Typhlops sp.
Ameiva wetmorei
Anolis cristatellus cristatellus Cayo Palominos (18.35°/-65.57°)
Anolis poncensis Ameiva exsul
Cyclura nubila (I) Anolis cristatellus wileyae
Iguana iguana (I) Anolis pulchellus
Sphaerodactylus nicholsi Sphaerodactylus townsendi
Typhlops hypomethes
Isla Mata Seca (17.96°/-67.01°) Typhlops platycephalus
Anolis cristatellus cristatellus
Cayo Piñeritos (18.24°/-65.59°)
Isla Morrillito (= Platillo) (17.88°/-66.53°)
Anolis cristatellus wileyae
Ameiva exsul
Ameiva wetmorei Cayo Santiago (18.16°/-65.73°)
Anolis cristatellus cristatellus Leptodactylus albilabris
Sphaerodactylus townsendi Rhinella marina (I)
Borikenophis portoricensis prymnus Ameiva exsul
Anolis cristatellus cristatellus
Isla Romero III (17.95°/-66.98°)
Anolis pulchellus
Anolis cristatellus cristatellus
Anolis stratulus
Long Island (= Long Key) (17.96°/-66.99°) Sphaerodactylus gaigeae
Anolis cristatellus cristatellus Sphaerodactylus macrolepis grandisquamis
Spiny Butte (17.96°/-66.99°) Borikenophis portoricensis portoricensis
Ameiva exsul Magliophis stahli
Anolis cristatellus cristatellus Isla Cabras (18.21°/-65.60°)
Sphaerodactylus nicholsi Ameiva exsul
Cays off the Eastern Coast of Puerto Rico Anolis cristatellus ssp.
Cabeza de Perro (= North Cabras Island) (18.25°/-65.58°) Sphaerodactylus townsendi
Ameiva exsul Isla Piñeros (18.25°/-65.59°)
Anolis cristatellus cristatellus Eleutherodactylus antillensis
Anolis pulchellus Leptodactylus albilabris
Sphaerodactylus macrolepis grandisquamis Ameiva exsul
Borikenophis portoricensis ssp. Amphisbaena caeca
Cayo Ahogado (18.32°/-65.62°) Anolis cristatellus wileyae
Ameiva exsul (W) Anolis pulchellus
Anolis cristatellus cristatellus (W) Anolis stratulus
Anolis pulchellus (W) Sphaerodactylus gaigeae
Cayo Algodones (18.19°/-65.68°) Sphaerodactylus macrolepis stibarus
Leptodactylus albilabris Sphaerodactylus townsendi
Rhinella marina (I) Borikenophis portoricensis ssp.
Ameiva exsul Typhlops hypomethes
Anolis cristatellus cristatellus x A. c. wileyae Isla de Ramos (18.31°/-65.61°)
Anolis pulchellus Leptodactylus albilabris
Sphaerodactylus macrolepis grandisquamis Ameiva exsul
Cayo Batata (18.12°/-65.77°) Anolis cristatellus cristatellus x A. c. wileyae
Ameiva exsul Anolis pulchellus
Anolis cristatellus cristatellus Sphaerodactylus townsendi
Anolis pulchellus Isleta Marina (= Cayo Zancudo) (18.34°/-65.62°)
Sphaerodactylus macrolepis grandisquamis Ameiva exsul
140 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Anolis cristatellus cristatellus x A. c. wileyae Sphaerodactylus townsendi

Anolis pulchellus Typhlops richardii
Levins Rock (18.18°/-65.71°) Vieques (= Bieque = Crab Island) (18.12°/-65.42°) (137.66 km2)
Ameiva exsul (W) Eleutherodactylus antillensis
Cayos de La Cordillera Eleutherodactylus cochranae
Eleutherodactylus coqui (I?)
La Blanquilla (18.37°/-65.55°)
Leptodactylus albilabris
Ameiva exsul
Lithobates catesbeianus (I)
Anolis cristatellus wileyae
Rhinella marina (I)
Sphaerodactylus townsendi
Trachemys stejnegeri stejnegeri (I?)
Booby Hatch Cay (= Booby Rock) (18.37°/-65.55°) Ameiva exsul
Anolis cristatellus wileyae Anolis cristatellus wileyae
Sphaerodactylus townsendi Anolis pulchellus
Cayo Diablo (= Cayo La Llave) (18.36°/-65.53°) Anolis roosevelti (PE)
Ameiva exsul Anolis stratulus
Anolis cristatellus wileyae Hemidactylus mabouia (I?)
Epicrates granti Sphaerodactylus gaigeae
Typhlops richardii Sphaerodactylus macrolepis inigoi
Sphaerodactylus roosevelti
Cayo Icacos (= Hicacos) (18.39°/-65.59°) Sphaerodactylus townsendi
Leptodactylus albilabris Spondylurus nitidus (PX)
Ameiva exsul Borikenophis portoricensis aphantus
Anolis cristatellus wileyae Typhlops hypomethes
Anolis pulchellus Typhlops platycephalus
Iguana iguana (I) Caiman crocodilus (W)
Sphaerodactylus townsendi
Spondylurus nitidus Cayo de Afuera (= Cayo Real) (18.09°/-65.47°)
Ameiva exsul
Cayo Lobos (18.38°/-65.57°) Amphisbaena caeca
Ameiva exsul Sphaerodactylus macrolepis inigoi
Anolis cristatellus wileyae Sphaerodactylus townsendi
Anolis pulchellus
Sphaerodactylus townsendi Cayo de Tierra (18.09°/-65.47°)
Ameiva exsul
Cayo Ratones (off Fajardo) (18.38°/-65.58°) Hemidactylus mabouia (I?)
Ameiva exsul Sphaerodactylus macrolepis inigoi
Anolis cristatellus wileyae Sphaerodactylus townsendi
Anolis pulchellus Typhlops sp.
Sphaerodactylus townsendi
Isla Chiva (18.11°/-65.38°)
Epicrates granti (I)
Anolis cristatellus wileyae
Grant Rock (18.37°/-65.55°) Sphaerodactylus townsendi
Anolis cristatellus wileyae
Culebra (18.31°/-65.28°) (27.28 km2)
Sphaerodactylus townsendi
Eleutherodactylus antillensis
Konyoki (18.38°/-65.58°) Eleutherodactylus cochranae
Ameiva exsul Eleutherodactylus coqui (I)
Anolis cristatellus wileyae Leptodactylus albilabris
Sphaerodactylus townsendi Rhinella marina (I)
MacKenzie Key (18.37°/-65.55°) Trachemys stejnegeri stejnegeri (I?)
Anolis cristatellus wileyae Ameiva exsul
Sphaerodactylus townsendi Anolis cristatellus wileyae
Anolis pulchellus
Pressick Rock (18.38°/-65.57°) Anolis roosevelti (PE)
Sphaerodactylus townsendi Anolis stratulus
Surprise Key (18.37°/-65.55°) Hemidactylus mabouia (I?)
Anolis cristatellus wileyae Iguana iguana (I)
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 141

Sphaerodactylus macrolepis macrolepis Chelonoidis carbonaria (I?)

Sphaerodactylus townsendi Trachemys scripta elegans (I)
Spondylurus culebrae Ameiva exsul
Spondylurus nitidus Amphisbaena fenestrata
Borikenophis portoricensis richardi Anolis leachii (I, E?)
Epicrates granti Anolis cristatellus wileyae
Magliophis exiguus exiguus Anolis pulchellus
Typhlops richardii Anolis stratulus
Cyclura pinguis (F*)
Cayo Botella (= Heatwole Island) (18.32°/-65.24°)
Gymnophthalmus underwoodi (I?)
Anolis cristatellus wileyae
Hemidactylus mabouia (I?)
Sphaerodactylus macrolepis macrolepis
Iguana iguana (I)
Cayo Geniqui East (= Palada Cays) (18.34°/-65.23°) Sphaerodactylus macrolepis macrolepis
Sphaerodactylus macrolepis macrolepis Spondylurus semitaeniatus (PX)
Cayo Luis Peña (= Southwest Key) (18.31°/-65.33°) Spondylurus sloanii (PX)
Anolis cristatellus wileyae Spondylurus spilonotus (PE)
Anolis pulchellus Thecadactylus rapicauda (PX)
Sphaerodactylus macrolepis macrolepis Boa constrictor ssp. (W)
Spondylurus nitidus Borikenophis portoricensis richardi (PX, I)
Borikenophis portoricensis richardi Epicrates granti
Typhlops richardii Magliophis exiguus exiguus
Pantherophis guttatus (I)
Cayo Norte (18.34°/-65.26°) Python regius (W)
Ameiva exsul Typhlops richardii
Anolis cristatellus wileyae
Anolis pulchellus Big Flat Cay (18.32°/-64.99°) (0.012 km2)
Sphaerodactylus macrolepis macrolepis Ameiva exsul
Spondylurus nitidus Anolis cristatellus wileyae
Typhlops richardii Sphaerodactylus macrolepis macrolepis
Bovoni Cay (18.31°/-64.87°) (0.20 km2)
Cayo Pirata (= Chicken Island) (18.31°/-65.30°)
Eleutherodactylus cochranae
Anolis cristatellus wileyae
Ameiva exsul
Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae
Culebrita (18.31°/-65.23°) Anolis stratulus
Ameiva exsul Hemidactylus mabouia (I?)
Anolis cristatellus wileyae Sphaerodactylus macrolepis macrolepis
Sphaerodactylus macrolepis macrolepis
Buck Island (western and larger of the Capella Islands)
Spondylurus culebrae
(18.28°/-64.89°) (0.17 km2)
Typhlops richardii
Anolis cristatellus wileyae
Isla Pela (= Bare Cactus Island) (18.30°/-65.25°) Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Spondylurus semitaeniatus
Sphaerodactylus macrolepis macrolepis Spondylurus sloanii
Pelaita (= Villa del Mar) (18.30°/-65.25°) Borikenophis portoricensis nicholsi
Anolis cristatellus wileyae Capella Island (= Eastern Island = unnamed island, eastern
Sphaerodactylus macrolepis macrolepis and smaller of the Capella Islands) (18.28°/-64.89°) (0.09 km2)
St. Thomas (18.35°/-64.94°) (70.23 km2) Anolis cristatellus wileyae
Eleutherodactylus antillensis Spondylurus semitaeniatus
Eleutherodactylus cochranae Spondylurus sloanii
Eleutherodactylus coqui (I) Cas Cay (18.31°/-64.87°) (0.06 km2)
Eleutherodactylus lentus Ameiva exsul
Eleutherodactylus schwartzi (?) Anolis cristatellus wileyae
Leptodactylus albilabris Anolis stratulus
Osteopilus septentrionalis (I) Iguana iguana (I)
Rhinella marina (I) Sphaerodactylus macrolepis macrolepis
142 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Cockroach Island (18.40°/-65.06°) (0.08 km2) Chelonoidis carbonaria (I?)

Anolis cristatellus wileyae Ameiva exsul
Sphaerodactylus macrolepis macrolepis Anolis cristatellus wileyae
Borikenophis portoricensis richardi Anolis pulchellus
Congo Cay (18.37°/-64.80°) (0.10 km2) Anolis stratulus
Anolis cristatellus wileyae Hemidactylus mabouia (I?)
Iguana iguana (I)
Anolis stratulus
Sphaerodactylus macrolepis macrolepis
Sphaerodactylus macrolepis macrolepis
Magliophis exiguus exiguus
Borikenophis portoricensis richardi
Typhlops richardii Inner Brass Island (18.38°/-64.97°) (0.52 km2)
Osteopilus septentrionalis (I)
Current Rock (18.31°/-64.83°) (0.002 km2)
Ameiva exsul
Anolis cristatellus wileyae
Anolis cristatellus wileyae
Dog Island (18.30°/-64.82°) (0.05 km2) Anolis stratulus
Ameiva exsul Sphaerodactylus macrolepis macrolepis (?)
Anolis cristatellus wileyae Borikenophis portoricensis ssp. (?)
Sphaerodactylus macrolepis macrolepis
Little Hans Lollik Island (18.41°/-64.91°) (0.41 km2)
Borikenophis portoricensis richardi
Ameiva exsul
Dutchcap Cay (= Dutchman’s Cap) (18.38°/-65.06°) (0.13 km2) Anolis cristatellus wileyae
Ameiva exsul Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Typhlops richardii
Sphaerodactylus macrolepis macrolepis
Little St. James Island (18.30°/-64.83°) (0.28 km2)
Borikenophis portoricensis richardi
Eleutherodactylus sp. (W)
Fish Cay (18.31°/-64.83°) (0.001 km2) Osteopilus septentrionalis (I)
Anolis cristatellus wileyae Ameiva exsul
Hemidactylus mabouia (I?) Anolis cristatellus wileyae
French Cap Cay (= Frenchman’s Cap) (18.23°/-64.85°) Anolis pulchellus
(0.04 km2) Anolis stratulus
Sphaerodactylus townsendi Iguana iguana (I)
Sphaerodactylus macrolepis macrolepis
Grass Cay (18.36°/-64.83°) (0.20 km2) Borikenophis portoricensis richardi
Anolis cristatellus wileyae Pantherophis guttatus (W)
Sphaerodactylus macrolepis macrolepis
Lovango Cay (18.36°/-64.81°) (0.48 km2)
Borikenophis portoricensis richardi
Chelonoidis carbonaria (I?)
Great St. James Island (18.31°/-64.83°) (0.63 km2) Ameiva exsul
Ameiva exsul Anolis cristatellus wileyae
Amphisbaena fenestrata Anolis pulchellus
Anolis cristatellus wileyae Anolis stratulus
Anolis pulchellus Hemidactylus mabouia (I?)
Anolis stratulus Borikenophis portoricensis richardi
Sphaerodactylus macrolepis macrolepis
Mingo Cay (18.36°/-64.82°) (0.20 km2)
Spondylurus sp.
Ameiva exsul
Borikenophis portoricensis richardi
Anolis cristatellus wileyae
Hans Lollik Island (18.40°/-64.91°) (1.98 km2) Anolis pulchellus
Ameiva exsul Anolis stratulus
Anolis cristatellus wileyae Sphaerodactylus macrolepis macrolepis
Anolis pulchellus Borikenophis portoricensis richardi
Anolis stratulus
Outer Brass Island (18.39°/-64.97°) (0.44 km2)
Sphaerodactylus macrolepis macrolepis Ameiva exsul
Borikenophis portoricensis richardi Anolis cristatellus wileyae
Hassel Island (18.33°/-64.94°) (0.56 km2) Anolis pulchellus
Eleutherodactylus cochranae Anolis stratulus
Eleutherodactylus lentus Sphaerodactylus macrolepis macrolepis
Osteopilus septentrionalis (I) Borikenophis portoricensis richardi
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 143

Patricia Cay (18.31°/-64.87°) (0.14 km2) West Cay (18.36°/-65.05°) (0.16 km2)
Ameiva exsul Ameiva exsul
Anolis cristatellus wileyae Anolis cristatellus wileyae
Anolis stratulus Anolis stratulus
Hemidactylus mabouia (I?) Sphaerodactylus macrolepis macrolepis
Iguana iguana (I) St. John (18.34°/-64.75°) (49.67 km2)
Sphaerodactylus macrolepis macrolepis Eleutherodactylus antillensis
Rotto Cay (18.31°/-64.86°) (0.008 km2) Eleutherodactylus cochranae
Ameiva exsul Eleutherodactylus coqui (I)
Anolis cristatellus wileyae Eleutherodactylus lentus
Sphaerodactylus macrolepis macrolepis Eleutherodactylus schwartzi (PX)
Saba Island (= Little Saba Island) (18.31°/-65.00°) (0.12 km2) Leptodactylus albilabris
Ameiva exsul Osteopilus septentrionalis (I)
Anolis cristatellus wileyae Chelonoidis carbonaria (I?)
Anolis stratulus Ameiva exsul
Sphaerodactylus macrolepis macrolepis Amphisbaena fenestrata
Spondylurus sloanii Anolis cristatellus wileyae
Borikenophis portoricensis richardi Anolis pulchellus
Anolis roosevelti (PE)
Sail Rock (18.29°/-65.10°) (0.007 km2) Anolis stratulus
Sphaerodactylus macrolepis macrolepis Hemidactylus mabouia (I?)
Salt Cay (18.36°/-65.05°) (0.23 km2) Iguana iguana (I)
Ameiva exsul Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Spondylurus spilonotus (PE)
Sphaerodactylus macrolepis macrolepis Borikenophis portoricensis richardi (PX)
Spondylurus sp. Magliophis exiguus exiguus
Savana Island (18.34°/-65.08°) (0.70 km2) Typhlops richardii
Ameiva exsul Cinnamon Cay (18.36°/-64.76°) (0.004 km2)
Anolis cristatellus wileyae Anolis cristatellus wileyae
Anolis stratulus Sphaerodactylus macrolepis macrolepis
Sphaerodactylus macrolepis macrolepis
Cocoloba Cay (18.32°/-64.76°) (0.004 km2)
Borikenophis portoricensis richardi
Anolis cristatellus wileyae
Shark Island (18.34°/-64.84°) (0.005 km2) Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae
Flanagan Island (18.33°/-64.65°) (0.09 km2)
Thatch Cay (18.36°/-64.86°) (0.96 km ) 2
Ameiva exsul
Eleutherodactylus cochranae Anolis cristatellus wileyae
Ameiva exsul Anolis stratulus
Anolis cristatellus wileyae Sphaerodactylus macrolepis macrolepis
Anolis stratulus Spondylurus sp.
Sphaerodactylus macrolepis macrolepis Typhlops richardii
Borikenophis portoricensis richardi
Henley Cay (one of the Durloe Cays) (18.35°/-64.79°)
Turtledove Cay (18.31°/-65.00) (.02 km2) (0.05 km2)
Anolis cristatellus wileyae Anolis cristatellus wileyae
Sphaerodactylus macrolepis macrolepis Hemidactylus mabouia (I?)
Water Island (18.32°/-64.95°) (1.99 km2) Sphaerodactylus macrolepis macrolepis
Chelonoidis carbonaria (I?) Leduck Island (18.32°/-64.69°) (0.05 km2)
Ameiva exsul Ameiva exsul
Anolis cristatellus wileyae Anolis cristatellus wileyae
Anolis stratulus Anolis stratulus
Hemidactylus mabouia (I?)
Ramgoat Cay (18.36°/-64.79°) (0.011 km2)
Iguana iguana (I)
Anolis cristatellus wileyae
Sphaerodactylus macrolepis macrolepis
Spondylurus sloanii (PX) Rata Cay (18.36°/-64.80°) (0.002 km2)
Borikenophis portoricensis richardi Anolis cristatellus wileyae
144 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Steven Cay (18.33°/-64.81°) (0.008 km2) Spondylurus sloanii

Anolis cristatellus wileyae Borikenophis portoricensis ssp.
Anolis stratulus Sandy Cay (18.43°/-64.72°) (0.06 km2)
Iguana iguana (I) Ameiva exsul
Sphaerodactylus macrolepis macrolepis Anolis cristatellus wileyae
Epicrates granti (I) Hemidactylus mabouia (I?)
Trunk Cay (18.36°/-64.77°) (0.009 km2) Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Sandy Spit (18.43°/-64.70°) (0.001 km2)
Anolis stratulus Anolis cristatellus wileyae
Waterlemon Cay (18.37°/-64.72°) (0.003 km2) Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Watson Rock (= Cable Rock = Shark Rock) (18.44°/-64.83°)
Sphaerodactylus macrolepis macrolepis Sphaerodactylus macrolepis macrolepis
Whistling Cay (18.37°/-64.75°) (0.08 km2) Tortola (18.45°/-64.60°) (54.44 km2)
Anolis cristatellus wileyae Eleutherodactylus antillensis
Anolis stratulus Eleutherodactylus cochranae
Iguana iguana (I) Eleutherodactylus johnstonei (I)
Sphaerodactylus macrolepis macrolepis Eleutherodactylus schwartzi
Jost van Dyke (18.47°/-64.75°) (8.40 km2) Leptodactylus albilabris
Eleutherodactylus antillensis Osteopilus septentrionalis (I)
Eleutherodactylus lentus (I?) Chelonoidis carbonaria (I?)
Eleutherodactylus schwartzi Pseudemys nelsoni (I)
Leptodactylus albilabris Trachemys scripta elegans (I)
Amphisbaena fenestrata Ameiva exsul
Anolis cristatellus wileyae Ameiva sp. (I?)
Anolis pulchellus Amphisbaena fenestrata
Anolis stratulus Anolis cristatellus wileyae
Hemidactylus mabouia (I?) Anolis pulchellus
Iguana iguana (W?) Anolis roosevelti (PE)
Sphaerodactylus macrolepis macrolepis Anolis stratulus
Borikenophis portoricensis ssp. (PX) Hemidactylus mabouia (I?)
Epicrates granti Iguana iguana (I)
Magliophis exiguus exiguus Sphaerodactylus macrolepis macrolepis
Typhlops richardii Sphaerodactylus parthenopion
Great Tobago (18.45°/-64.80°) (0.87 km2) Spondylurus semitaeniatus
Ameiva exsul Borikenophis portoricensis richardi
Anolis cristatellus wileyae Epicrates granti
Hemidactylus mabouia (I?) Magliophis exiguus exiguus
Sphaerodactylus macrolepis macrolepis Pantherophis guttatus (W?)
Typhlops richardii
Green Cay (18.45°/-64.70°) (0.07 km2)
Anolis cristatellus wileyae Beef Island (18.45°/-64.52°) (3.72 km2)
Sphaerodactylus macrolepis macrolepis Eleutherodactylus antillensis
Borikenophis portoricensis ssp. Eleutherodactylus schwartzi
Leptodactylus albilabris
Little Jost van Dyke (18.45°/-64.72°) (0.63 km2) Osteopilus septentrionalis (I)
Ameiva exsul Ameiva exsul
Anolis cristatellus wileyae Anolis cristatellus wileyae
Anolis pulchellus Anolis pulchellus
Sphaerodactylus macrolepis macrolepis Anolis stratulus
Borikenophis portoricensis ssp. Hemidactylus mabouia (I?)
Typhlops richardii Sphaerodactylus macrolepis macrolepis
Little Tobago (18.43°/-64.85°) (0.22 km2) Borikenophis portoricensis anagadae
Anolis cristatellus wileyae Magliophis exiguus exiguus
Sphaerodactylus macrolepis macrolepis Typhlops richardii
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 145

Bellamy Cay (18.43°/-64.53°) (0.007 km2) Borikenophis portoricensis anegadae

Anolis cristatellus wileyae Epicrates granti (?)
Hemidactylus mabouia (I?) Magliophis exiguus exiguus
Sphaerodactylus macrolepis macrolepis Typhlops richardii
Buck Island (18.42°/-64.55°) (0.17 km2) Little Camanoe (18.45°/-64.53°) (0.16 km2)
Ameiva exsul Ameiva exsul
Anolis cristatellus wileyae Anolis cristatellus wileyae
Sphaerodactylus macrolepis macrolepis Sphaerodactylus macrolepis macrolepis
Frenchman’s Cay (18.38°/-64.70°) (0.24 km2) Little Thatch Island (18.38°/-64.72°) (0.24 km2)
Eleutherodactylus antillensis Eleutherodactylus schwartzi (I)
Eleutherodactylus schwartzi Anolis cristatellus wileyae
Ameiva exsul Anolis stratulus
Amphisbaena fenestrata Cyclura pinguis (I)
Anolis cristatellus wileyae Sphaerodactylus macrolepis macrolepis
Anolis pulchellus Spondylurus semitaeniatus
Anolis stratulus
Hemidactylus mabouia (I?) Marina Cay (18.45°/-64.52°) (0.010 km2)
Sphaerodactylus macrolepis macrolepis Ameiva exsul
Magliophis exiguus exiguus Anolis cristatellus wileyae
Anolis stratulus
Great Camanoe (18.48°/-64.53°) (3.37 km2) Hemidactylus mabouia (I?)
Eleutherodactylus antillensis
Sphaerodactylus macrolepis macrolepis
Ameiva exsul
Typhlops richardii
Amphisbaena fenestrata
Anolis cristatellus wileyae Scrub Island (18.47°/-64.52°) (0.97 km2)
Anolis pulchellus Ameiva exsul
Anolis stratulus Anolis cristatellus wileyae
Hemidactylus mabouia (I?) Anolis stratulus
Sphaerodactylus macrolepis macrolepis Sphaerodactylus macrolepis macrolepis
Spondylurus semitaeniatus Spondylurus sp.
Borikenophis portoricensis anegadae Peter Island (18.37°/-64.58°) (4.29 km2)
Epicrates granti Eleutherodactylus sp.
Magliophis exiguus exiguus Osteopilus septentrionalis (I)
Great Thatch Island (18.38°/-64.72°) (1.23 km2) Chelonoidis carbonaria (I?)
Eleutherodactylus antillensis Ameiva exsul
Eleutherodactylus cochranae Anolis cristatellus wileyae
Amphisbaena fenestrata Anolis pulchellus
Anolis cristatellus wileyae Anolis stratulus
Anolis pulchellus Hemidactylus mabouia (I?)
Anolis stratulus Iguana iguana (I)
Sphaerodactylus macrolepis macrolepis Sphaerodactylus macrolepis macrolepis
Guana Island (18.48°/-64.57°) (2.97 km2) Spondylurus sloanii
Eleutherodactylus antillensis Borikenophis portoricensis richardi
Osteopilus septentrionalis (W) Magliophis exiguus exiguus
Centrochelys sulcata (I) Pantherophis guttatus (W)
Chelonoidis carbonaria (I) Carrot Rock (18.32°/-64.57°) (0.013 km2)
Ameiva exsul Anolis ernestwilliamsi
Amphisbaena fenestrata Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Spondylurus macleani
Anolis pulchellus
Anolis stratulus Deadman’s Chest (= Dead Chest) (18.37°/-64.57°) (0.14 km2)
Cyclura pinguis (I) Ameiva exsul
Hemidactylus mabouia (I?) Anolis cristatellus wileyae
Sphaerodactylus macrolepis macrolepis Sphaerodactylus macrolepis macrolepis
Spondylurus semitaeniatus Spondylurus sp.
146 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Norman Island (18.33°/-64.62°) (2.57 km2) Fallen Jerusalem (18.42°/-64.45°) (0.12 km2)
Ameiva exsul Anolis cristatellus wileyae
Anolis cristatellus wileyae Anolis pulchellus
Anolis pulchellus Anolis stratulus
Anolis stratulus Sphaerodactylus macrolepis macrolepis
Cyclura pinguis (I) Spondylurus semitaeniatus
Sphaerodactylus macrolepis macrolepis
George Dog (18.5°/-64.45°) (0.15 km2)
Spondylurus sloanii
Ameiva exsul
Borikenophis portoricensis ssp.
Anolis cristatellus wileyae
Typhlops richardii
Sphaerodactylus macrolepis macrolepis
Pelican Island (18.33°/-64.62°) (0.03 km2)
Ginger Island (18.40°/-64.47°) (1.05 km2)
Anolis cristatellus wileyae
Ameiva exsul
Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae
Skipper Cay (= Key Cay) (18.34°/-64.60°) (0.011 km2) Anolis stratulus
Anolis cristatellus wileyae Sphaerodactylus macrolepis macrolepis
Virgin Gorda (18.50°/-64.40°) (21.30 km2) Spondylurus semitaeniatus
Eleutherodactylus antillensis Borikenophis portoricensis ssp.
Eleutherodactylus cochranae Great Dog (18.48°/-64.45°) (0.33 km2)
Eleutherodactylus schwartzi Eleutherodactylus schwartzi
Osteopilus septentrionalis (I) Ameiva exsul
Peltophryne lemur (PX) Anolis cristatellus wileyae
Chelonoidis carbonaria (I?) Sphaerodactylus macrolepis macrolepis
Ameiva exsul Typhlops sp.
Amphisbaena fenestrata
Anolis cristatellus wileyae Moskito Island (= Mosquito Island) (18.50°/-64.38°) (0.50 km2)
Anolis pulchellus Osteopilus septentrionalis (I)
Anolis stratulus Ameiva exsul
Hemidactylus mabouia (I?) Anolis cristatellus wileyae
Iguana iguana (I) Anolis stratulus
Sphaerodactylus macrolepis macrolepis Cyclura pinguis (I)
Sphaerodactylus parthenopion Hemidactylus mabouia (I?)
Spondylurus semitaeniatus Sphaerodactylus macrolepis macrolepis
Borikenophis portoricensis anegadae Sphaerodactylus parthenopion
Magliophis exiguus exiguus Spondylurus semitaeniatus
Typhlops naugus Thecadactylus rapicauda
Borikenophis portoricensis anegadae
Broken Jerusalem (18.40°/-64.45°) (0.009 km2)
Anolis cristatellus wileyae Necker Island (18.55°/-64.35°) (0.30 km2)
Eleutherodactylus antillensis (I)
Cistern Rock (18.38°/-64.50°) (0.001 km2)
Osteopilus septentrionalis (W)
Anolis cristatellus wileyae
Centrochelys sulcata (I)
Cooper Island (18.37°/-64.50°) (1.38 km2) Chelonoidis carbonaria (I)
Ameiva exsul Ameiva exsul
Anolis cristatellus wileyae Anolis cristatellus wileyae
Anolis stratulus Anolis pulchellus
Hemidactylus mabouia (I?) Anolis stratulus
Sphaerodactylus macrolepis macrolepis Cyclura pinguis (I)
Borikenophis portoricensis ssp. Sphaerodactylus macrolepis macrolepis
East Seal Dog (18.50°/-64.43°) (0.008 km2) Spondylurus semitaeniatus
Anolis cristatellus wileyae Thecadactylus rapicauda
Sphaerodactylus macrolepis macrolepis Borikenophis portoricensis anegadae
Typhlops naugus
Eustatia Island (18.52°/-64.37°) (0.11 km2)
Ameiva exsul Prickly Pear Island (18.50°/-64.37°) (0.70 km2)
Anolis cristatellus wileyae Ameiva exsul
Sphaerodactylus macrolepis macrolepis Anolis cristatellus wileyae
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 147

Anolis stratulus Osteopilus septentrionalis (I)

Sphaerodactylus macrolepis macrolepis Lithobates catesbeianus (W)
Spondylurus sp. Lithobates pipiens (W)
Typhlops naugus Rhinella marina (I)
Round Rock (18.40°/-64.45°) (0.08 km2) Chelonoidis carbonaria (I?)
Ameiva exsul Trachemys scripta elegans (I)
Anolis cristatellus wileyae Ameiva exsul (I)
Sphaerodactylus macrolepis macrolepis Ameiva polops (PX)
Spondylurus semitaeniatus Anolis acutus
Capitellum parvicruzae (PE)
Saba Rock (18.50°/-64.45°) (0.002 km2) Hemidactylus mabouia (I?)
Anolis cristatellus wileyae Iguana iguana (I)
Sphaerodactylus macrolepis macrolepis Sphaerodactylus beattyi beattyi
Salt Island (18.38°/-64.52°) (0.78 km2) Sphaerodactylus beattyi seamani
Ameiva exsul Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae Spondylurus magnacruzae (PE)
Anolis stratulus Thecadactylus rapicauda
Hemidactylus mabouia (I?) Borikenophis portoricensis (W)
Sphaerodactylus macrolepis macrolepis Borikenophis sancticrucis (PE)
Spondylurus semitaeniatus Pantherophis guttatus (W)
Spondylurus sloanii Ramphotyphlops braminus (I)
Borikenophis portoricensis richardi Thamnophis sp. (W)
Typhlops richardii
South Cockroach (= Big Cockroach = Cockroach Island)
(18.48°/-64.45°) (0.004 km2) Buck Island (17.79°/-64.62°) (0.73 km2)
Anolis cristatellus wileyae Ameiva polops (I)
Sphaerodactylus macrolepis macrolepis Anolis acutus
Sphaerodactylus beattyi beattyi
West Dog (18.48°/-64.47°) (0.13 km2)
Sphaerodactylus macrolepis macrolepis
Anolis cristatellus wileyae
Sphaerodactylus macrolepis macrolepis Green Cay (17.77°/-64.67°) (0.05 km2)
Ameiva polops
West Seal Dog (18.50°/-64.43°) (0.02 km2)
Anolis acutus
Anolis cristatellus wileyae
Sphaerodactylus beattyi beattyi
Sphaerodactylus macrolepis macrolepis
Sphaerodactylus macrolepis macrolepis
Anegada (18.75°/-64.33°) (38.72 km2) Spondylurus magnacruzae (PE)
Leptodactylus albilabris Borikenophis sancticrucis (PE)
Ameiva exsul
Protestant Cay (17.75°/-64.70°) (0.02 km2)
Anolis cristatellus wileyae
Ameiva polops
Anolis pulchellus
Anolis acutus
Anolis stratulus
Sphaerodactylus macrolepis macrolepis
Cyclura pinguis
Hemidactylus mabouia (I?) Ruth Island (17.68°/-64.76°) (0.08 km2)
Sphaerodactylus macrolepis macrolepis Ameiva polops (I)
Spondylurus anegadae
Thecadactylus rapicauda
Borikenophis portoricensis anegadae
Magliophis exiguus exiguus
Typhlops catapontus

ST. CROIX BANK

St. Croix (17.74°/-64.74°) (211.78 km2)
Eleutherodactylus antillensis (I?)
Eleutherodactylus coqui (I)
Eleutherodactylus lentus
Eleutherodactylus schwartzi (I?)
Leptodactylus albilabris
148 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

LESSER ANTILLES

Robert W. Henderson1 and Michel Breuil2

About 120 species of amphibians and reptiles Osteopilus septentrionalis (I)
have been documented on 139 islands in the Rhinella marina (I)
Chelonoidis carbonaria (F*, I?)
Lesser Antilles. In addition to the sources noted Phrynops geoffroanus (W)
in the Introduction and our own fieldwork, a Ameiva plei plei
useful paper on the herpetofauna of the Windward Anolis carolinensis (I)
Islands by Corke (1992) and recent books on the Anolis gingivinus
Anolis pogus (PE)
herpetofauna of the Eastern Caribbean (Malhotra
Hemidactylus mabouia (I?)
& Thorpe 1999), the Guadeloupean Archipelago Iguana delicatissima
(Breuil 2002), Anguilla (Hodge et al. 2003), and Iguana iguana (I)
the Dutch Windward Islands (Powell et al. 2005) Leiocephalus cf. cuneus (F)
were consulted. In addition, a recent collection Sphaerodactylus parvus
Sphaerodactylus sputator
of papers concerned with the herpetofaunas of Spondylurus powelli
Anguilla (Hodge et al. 2011), the French West Thecadactylus rapicauda
Indies (Lorvelec et al. 2011), Antigua, Barbuda, Alsophis rijgersmaei
and Redonda (Daltry 2011), the Dutch Windward Pantherophis guttatus (W)
Islands (Powell 2011), Dominica (Malhotra et Ramphotyphlops braminus (I)
al. 2011), Guadeloupe and Martinique (Breuil Anguillita (18.16°/-63.18°) (0.05 km2)
Anolis gingivinus
2011), St. Vincent (Powell & Henderson 2011),
Anolis pogus (I, E)
the Grenadine Islands (Daudin & de Silva 2011),
Dog Island (18.28°/-63.25°) (2.06 km2)
the Grenada Bank (Henderson & Berg 2011), and Ameiva plei plei
Barbados (Fields & Horrocks 2011) have been Anolis gingivinus
especially helpful. Information on fossil frogs and Sphaerodactylus parvus
lizards was taken from Pregill et al. (1994). Latitude Sphaerodactylus sputator
and longitude were gleaned from Google Earth®. Spondylurus powelli
Little Scrub (18.30°/-62.96°) (0.05 km2)
ACKNOWLEDGMENT Ameiva corax
Sphaerodactylus sputator

We thank Mark de Silva for information regarding Prickly Pear Cay East (18.26°/-63.17°) (0.33 km2)
Ameiva plei
islets associated with St. Vincent and the Anolis gingivinus
Grenadines.
Prickly Pear Cay West (18.27°/-63.19°) (0.34 km2)
Anolis gingivinus
SOMBRERO BANK
Sombrero (18.36°/-63.03°) (0.37 km2) Scilly Cay (18.26°/-63.00°) (0.007 km2)
Chelonoidis sombrerensis (F) Ameiva plei plei
Anolis gingivinus
Ameiva corvina
Anolis gingivinus Scrub Island (18.29°/-62.95°) (3.45 km2)
Sphaerodactylus sp. Ameiva plei plei
Anolis gingivinus
ANGUILLA BANK Sphaerodactylus parvus
Anguilla (18.23°/-63.03°) (90.7 km2) Sphaerodactylus sputator
Eleutherodactylus johnstonei (I) Alsophis rijgersmaei
1
Section of Vertebrate Zoology, Milwaukee Public Museum, Milwaukee, Wisconsin 53233, USA <[email protected]>
2
Département de Systématique et d’Évolution, Taxonomie et Collections, Reptiles et Amphibiens, Muséum national d’Histoire naturelle, 75005 Paris
<[email protected]>
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 149

Figure 9. Map of the Lesser Antilles.

150 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

West Cay (off Dog Is.) (18.28°/-63.27°) (0.02 km2) L’Îlet au Vent (17.95°/-62.88°) (0.01 km2)
Ameiva plei Ameiva plei plei
Saint-Barthélemy (= St. Barts) (17.89°/-62.83°) (22 km2) Anolis gingivinus
Eleutherodactylus johnstonei (I) Iguana delicatissima (W)
Eleutherodactylus martinicensis (I) La Petite Islette (17.95°/-62.90°) (0.01 km2)
Eleutherodactylus sp. (PE) Ameiva plei plei
Osteopilus septentrionalis (I) Anolis gingivinus
Pelophylax kl. esculentus (I) Iguana delicatissima (W)
Salamandra salamandra (W) Saint-Martin/Sint Maarten (18.06°/-63.05°) (85 km2)
Chelonoidis carbonaria (I?) Eleutherodactylus johnstonei (I)
Trachemys scripta elegans (W) Eleutherodactylus martinicensis (I)
Ameiva plei plei Eleutherodactylus sp. (PE)
Anolis gingivinus Osteopilus septentrionalis (I)
Anolis pogus (PX) Chelonoidis carbonaria (I?)
Hemidactylus mabouia (I?) Trachemys scripta elegans (I)
Iguana delicatissima Ameiva plei analifera
Iguana iguana (I) Ameiva plei plei
Iguana iguana x I. delicatissima Gymnophthalmus underwoodi (I)
Sphaerodactylus parvus Anolis bimaculatus (I, E?)
Sphaerodactylus sputator Anolis cristatellus (I)
Spondylurus powelli Anolis gingivinus
Thecadactylus oskrobapreinorum (?) Anolis pogus
Thecadactylus rapicauda Anolis sagrei ssp. (I)
Alsophis rijgersmaei Hemidactylus mabouia (I?)
Pantherophis guttatus (W) Iguana delicatissima (PX)
Python bivittatus (W) Iguana iguana (I)
Python regius (W) Sphaerodactylus parvus
Thamnophis sirtalis ssp. (W) Sphaerodactylus sputator
Ramphotyphlops braminus (I) Spondylurus martinae (PE)
Typhlops annae Thecadactylus oskrobapreinorum
Île Chevreau ou Île Bonhomme (17.93°/-62.85°) (0.25 km2) Thecadactylus rapicauda
Ameiva plei plei Alsophis rijgersmaei (PX)
Anolis gingivinus Alsophis rufiventris (W)
Iguana delicatissima (PX) Boa constrictor ssp. (W)
Epicrates cenchria (W)
Île Coco (17.87°/-62.81°) (0.04 km2)
Pantherophis guttatus (W)
Anolis gingivinus
Python curtus group (W)
Île Fourchue (17.96°/-62.90°) (0.30 km2) Python regius (W)
Ameiva plei plei Ramphotyphlops braminus (I)
Anolis gingivinus
Guana Cay off Pelikan (18.02°/-63.02°) (0.02 km2)
Iguana delicatissima
Anolis gingivinus
Sphaerodactylus sputator
Pantherophis guttatus (I?)
Alsophis rijgersmaei (PX)
Îlet Pinel (18.11°/-63.01°) (0.05 km2)
Île Frégate (17.94°/-62.84°) (0.11 km2)
Chelonoidis carbonaria (W)
Ameiva plei plei
Anolis gingivinus
Anolis gingivinus
Sphaerodactylus parvus
Hemidactylus mabouia (W)
Sphaerodactylus sputator
Iguana delicatissima (E, I)
Iguana iguana (I)
Île Toc Vers (17.94°/-62.82°) (0.02 km2)
Tintamarre (18.12°/-62.98°) (0.50 km2)
Ameiva plei plei
Chelonoidis carbonaria (I?)
Anolis gingivinus
Ameiva plei plei
Iguana delicatissima
Anolis gingivinus
Îlet Tortue (17.92°/-62.80) Sphaerodactylus parvus
Alsophis rijgersmaei Iguana iguana (I, from St.-Martin)
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 151

SABA BANK Sphaerodactylus sabanus

Saba (17.63°/-63.23°) (13 km )
2 Sphaerodactylus sputator
Eleutherodactylus johnstonei (I) Thecadactylus rapicauda
Osteopilus septentrionalis (W) Alsophis rufiventris
Chelonoidis carbonaria (W) Ramphotyphlops braminus (I)
Anolis sabanus
Hemidactylus mabouia (I?) ANTIGUA BANK
Iguana iguana Antigua (17.08°/-61.80°) (280 km2)
Sphaerodactylus sabanus Eleutherodactylus johnstonei
Thecadactylus rapicauda Eleutherodactylus martinicensis (I?)
Alsophis rufiventris Osteopilus septentrionalis (I)
Rhinella marina (I)
ST. CHRISTOPHER BANK Chelonoidis carbonaria (I?)
St. Christopher (= St. Kitts) (17.36°/-62.78°) (170 km2) Ameiva griswoldi
Eleutherodactylus johnstonei (I) Anolis leachii
Leptodactylus fallax (PX) Anolis wattsi wattsi
Rhinella marina (I) Gymnophthalmus underwoodi (I)
Chelonoidis carbonaria (I?) Hemidactylus mabouia (I?)
Chelonoidis sp. (I?) Iguana delicatissima (PX)
Ameiva erythrocephala Iguana iguana (W)
Anolis bimaculatus Leiocephalus cuneus (F)
Anolis schwartzi Sphaerodactylus elegantulus
Gymnophthalmus underwoodi (I) Thecadactylus rapicauda
Hemidactylus mabouia (I?) Alsophis antiguae antiguae (PX)
Iguana delicatissima (F*) Boa sp. (F)
Sphaerodactylus sabanus Pantherophis guttatus (W)
Sphaerodactylus sputator Typhlops geotomus
Thecadactylus rapicauda
Alsophis rufiventris (PE) Barbuda (17.65°/-61.81°) (160 km2)
Ramphotyphlops braminus (I) Eleutherodactylus johnstonei
Typhlops geotomus Chelonoidis carbonaria (I?)
Ameiva griswoldi
Nevis (17.16°/-62.59°) (130 km2) Anolis leachii
Eleutherodactylus johnstonei (I) Anolis wattsi foresti
Osteopilus septentrionalis (I)
Gymnophthalmus underwoodi (I?)
Rhinella marina (I)
Iguana iguana (W)
Ameiva erythrocephala
Leiocephalus cuneus (F)
Anolis bimaculatus
Sphaerodactylus elegantulus
Anolis schwartzi
Thecadactylus rapicauda
Hemidactylus mabouia (I?)
Hemidactylus mabouia (PX)
Iguana delicatissima (PX)
Alsophis sp. (F)
Sphaerodactylus sabanus
Clelia cf. clelia (F)
Sphaerodactylus sputator
Typhlops geotomus
Thecadactylus rapicauda
Alsophis rufiventris (PE) Great Bird Island (17.14°/-61.72°) (0.09 km2)
Typhlops geotomus Ameiva griswoldi
St. Eustatius (17.49°/-62.98°) (20 km2) Anolis leachii
Eleutherodactylus johnstonei (I) Anolis wattsi wattsi
Chelonoidis carbonaria (W) Sphaerodactylus elegantulus
Chelonoidis sp. (PE) Alsophis antiguae sajdaki
Trachemys scripta elegans (W) Typhlops geotomus
Ameiva erythrocephala Green Island (17.07°/-61.67°) (0.43 km2)
Anolis bimaculatus Eleutherodactylus johnstonei
Anolis schwartzi Chelonoidis carbonaria (I)
Hemidactylus mabouia (I?) Ameiva griswoldi
Iguana delicatissima Anolis leachii
152 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Anolis wattsi wattsi Kinixys erosa (I, E)

Thecadactylus rapicaudus Kinixys homeana (I, E)
Alsophis antiguae sajdaki (I) Pelusios castaneus (W)
Guana Island (17.12°/-61.75) Trachemys scripta elegans (W)
Anolis wattsi wattsi Trachemys stejnegeri stejnegeri (I)
Ameiva cineracea (PE)
Long Island (17.15°/-61.75) Anolis marmoratus marmoratus
Ameiva griswoldi Anolis marmoratus alliaceus
Anolis leachii Anolis marmoratus girafus
Anolis wattsi wattsi Anolis marmoratus setosus
Rabbit Island (17.13°/-61.73°) (0.02 km2) Anolis marmoratus speciosus
Anolis leachii Gymnophthalmus underwoodi (I)
Anolis wattsi wattsi Hemidactylus mabouia (I?)
Alsophis antiguae sajdaki (I) Iguana delicatissima
Iguana iguana (I)
York (17.03°/-61.67)
Iguana iguana x Iguana delicatissima
Anolis leachii
Leiocephalus cf. L. cuneus (F)
Anolis wattsi wattsi
Diploglossus sp. (PX)
Mabuya guadeloupae (PE)
REDONDA BANK
Sphaerodactylus fantasticus fantasticus
Redonda (16.94°/-62.35)
Sphaerodactylus fantasticus orescius
Ameiva atrata
Thecadactylus rapicauda
Anolis nubilis
Alsophis antillensis
Copeoglossum redondae (PE)
Clelia sp. (PE)
Hemidactylus mabouia (I?)
Liophis juliae
Sphaerodactylus sp.
Morelia amethistina (W)
MONTSERRAT BANK Ramphotyphlops braminus (I)
Montserrat (16.74°/-62.19°) (100 km2) Typhlops guadeloupensis
Eleutherodactylus johnstonei (I) Guadeloupe (Grande-Terre) (16.33°/-61.46°) (590 km2)
Leptodactylus fallax Eleutherodactylus johnstonei (I)
Rhinella marina (I) Eleutherodactylus martinicensis
Chelonoidis carbonaria (I?) Leptodactylus fallax (I, E)
Ameiva pluvianotata Rhinella marina (I)
Anolis lividus Scinax cf. x-signatus (I)
Diploglossus montisserrati Chelonoidis carbonaria (I?)
Hemidactylus mabouia (I?) Pelusios castaneus (I)
Iguana iguana Trachemys scripta elegans (W)
Mabuya montserratae (PE) Trachemys stejnegeri stejnegeri (I)
Sphaerodactylus fantasticus ligniservulus Ameiva cineracea (F)
Thecadactylus rapicauda Anolis marmoratus inornatus
Alsophis manselli Anolis marmoratus speciosus
Typhlops monastus Gymnophthalmus pleii (?)
Gymnophthalmus underwoodi (I)
GUADELOUPE BANK Hemidactylus mabouia (I?)
Guadeloupe (Basse-Terre) (16.18°/-61.68°) (848 km2) Iguana delicatissima (PX)
Eleutherodactylus barlagnei Iguana iguana (I)
Eleutherodactylus johnstonei (I) Iguana iguana x I. delicatissima
Eleutherodactylus martinicensis Leiocephalus cf. L. cuneus (F)
Eleutherodactylus pinchoni Lepidodactylus lugubris (I)
Eleutherodactylus cf. planirostris (W) Mabuya grandisterrae (PE)
Leptodactylus fallax (I, E) Sphaerodactylus fantasticus karukera
Rhinella marina (I) Sphaerodactylus fantasticus tartaropylorus
Scinax cf. x-signatus (I) Thecadactylus rapicauda
Chelonoidis carbonaria (I?) Geckko gecko (I)
Chelonoidis denticulata (W) Alsophis antillensis
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 153

Liophis juliae Îlets de Pigeon, Îlet à Goyaves, Sud (= Petit Îlet) (16.66°/-
Ramphotyphlops braminus (I) 61.789°) (0.01 km2)
Typhlops guadeloupensis Anolis marmoratus girafus
Islets in the Grand Cul de Sac (between Grande- Iguana iguana
Terre and Basse-Terre, North) Iguana delicatissima (PX)
Îlet Macou (16.35°/-61.53°) (0.05 km2) Sphaerodactylus fantasticus ssp.
Anolis marmoratus inornatus La Désirade (16.32°/-61.06°) (22 km2)
Îlet Fajou (16.35°/-61.58°) (1.04 km2) Eleutherodactylus johnstonei (I)
Anolis marmoratus speciosus x Anolis m. setosus Eleutherodactylus martinicensis
Hemidactylus mabouia (I?) Scinax cf. x-signatus (I)
Sphaerodactylus fantasticus ssp. Ameiva sp. (F)
Anolis marmoratus desiradei
Îlet à Kahouanne (16.37°/-61.78°) (0.18 km2) Gymnophthalmus underwoodi (I)
Eleutherodactylus martinicensis Hemidactylus mabouia (I?)
Anolis kahouannensis Iguana delicatissima
Iguana delicatissima and°/or Iguana iguana Leiocephalus sp. (F)
Sphaerodactylus fantasticus ssp. Mabuya desiradae
Hemidactylus mabouia (I?) Sphaerodactylus fantasticus hippomanes
Thecadactylus rapicauda Thecadactylus rapicauda
Tête à l’Anglais (16.39°/-61.77°) (0.02 km2) Îles de la Petite Terre
Anolis kahouannensis Terre de Bas (16.17°/-61.12)
Hemidactylus mabouia (I?) Eleutherodactylus sp. (W)
Islets in the Petit Cul de Sac (between Grande-Terre Chelonoidis carbonaria (W)
and B asse -T erre , S outh ) Ameiva major (PE)
Grand Îlet (off Petit-Bourg) (16.18°/-61.58°) (0.005 km2) Anolis marmoratus chrysops
Ameiva cineracea (PE) Hemidactylus mabouia (I?)
Mabuya desiradae
Îlet Boissard (16.22°/-61.55°) (0.05 km2)
Sphaerodactylus fantasticus karukera
Anolis marmoratus ssp.
Thecadactylus rapicauda
Iguana iguana
Boa nebulosa (W)
Alsophis or Liophis
Terre de Haut (16.18°/-61.11)
Îlet Christophe (16.28°/-61.55°) (0.01 km2)
Anolis marmoratus chrysops
Anolis marmoratus speciosus
Sphaerodactylus fantasticus karukera
Îlet Fortune (16.15°/-61.55°) (0.01 km2) Hemidactylus mabouia (I?)
Sphaerodactylus fantasticus karukera x S. f. orescius Iguana delicatissima
Îlet Gosier (16.20°/-61.49°) (0.02 km2)
Anolis marmoratus speciosus ÎLE DES SAINTES BANK
Sphaerodactylus fantasticus karukera Terre-de-Bas (15.86°/-61.68°) (9.5 km2)
Eleutherodactylus johnstonei (I)
Îlet sans nom (16.22°/-61.53°) (0.02 km2) Eleutherodactylus martinicensis
Iguana iguana Chelonoidis carbonaria (I?)
Îlet-à-Cochons (16.21°/-61.54°) (0.20 km2) Trachemys scripta elegans (I?)
Eleutherodactylus martinicensis Trachemys stejnegeri stejnegeri (I)
Anolis marmoratus speciosus Anolis terraealtae caryae
Mabuya cochonae Hemidactylus mabouia (I?)
Iguana delicatissima (PX)
Islets west of Basse-Terre
Iguana iguana x I. delicatissima
Îlets de Pigeon, Îlet à Goyaves, Nord (= Grand Îlet) (16.66°/-
Iguana iguana (I)
61.789°) (0.05 km2)
Sphaerodactylus phyzacinus
Anolis marmoratus girafus
Thecadactylus rapicauda
Hemidactylus mabouia (I?)
Alsophis sanctonum danforthi
Iguana delicatissima (PX)
Iguana iguana Terre-de-Haut (15.87°/-61.58°) (4.5 km2)
Gymnophthalmus underwoodi (I) Eleutherodactylus johnstonei (I)
Sphaerodactylus fantasticus ssp. Eleutherodactylus martinicensis
154 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Chelonoidis carbonaria (I?) Eleutherodactylus martinicensis

Trachemys stejnegeri stejnegeri (I) Leptodactylus fallax
Anolis terraealtae terraealtae Osteopilus septentrionalis (W)
Gymnophthalmus underwoodi (I) Rhinella marina (W)
Hemidactylus mabouia (I?) Chelonoidis carbonaria (I?)
Thecadactylus rapicauda Trachemys stejnegeri stejnegeri (I, E)
Iguana delicatissima (PX) Ameiva fuscata
Iguana iguana (I) Anolis bimaculatus (W)
Iguana iguana x I. delicatissima Anolis cristatellus (I)
Sphaerodactylus phyzacinus Anolis oculatus oculatus
Alsophis sanctonum sanctonum Anolis oculatus cabritensis
Grande Îlet (= Mare Basse) (15.84°/-61.59°) (0.7 km2) Anolis oculatus montanus
Eleutherodactylus martinicensis Anolis oculatus winstoni
Chelonoidis carbonaria (I?) Gonatodes vittatus (W)
Pelusios castaneus (W) Gymnophthalmus pleii
Anolis terraealtae terraealtae Gymnophthalmus underwoodi (I)
Hemidactylus mabouia (I?) Hemidactylus mabouia (I?)
Iguana iguana Iguana delicatissima
Sphaerodactylus phyzacinus Mabuya dominicana
Sphaerodactylus fantasticus fuga
Îlet à Cabrit (15.88°/-61.59°) (0.4 km2) Sphaerodactylus microlepis (W)
Eleutherodactylus martinicensis Sphaerodactylus vincenti monilifer
Anolis terraealtae terraealtae Thecadactylus rapicauda
Hemidactylus mabouia (I?) Alsophis sibonius
Iguana iguana (I) Boa nebulosa
Sphaerodactylus phyzacinus Liophis juliae
Alsophis sanctonum ssp. (probably sanctonum) Typhlops dominicanus
La Coche (15.84°/-61.61°) (0.04 km2)
Anolis terraealtae terraealtae MARTINIQUE BANK
Hemidactylus mabouia (I?) Martinique (14.68°/-61.01°) (1,100 km2)
Iguana iguana Allobates chalcopis (I?)
Sphaerodactylus phyzacinus Eleutherodactylus johnstonei (I)
Eleutherodactylus martinicensis
MARIE GALANTE BANK Leptodactylus fallax (I, E)
Marie Galante (15.94°/-61.26°) (160 km2) Rhinella marina (I)
Eleutherodactylus johnstonei (I) Scinax ruber (I)
Eleutherodactylus martinicensis Scinax cf. x-signatus (I)
Scinax cf. x-signatus (I) Centrochelys sulcata (W)
Trachemys scripta elegans (W) Chelonoidis carbonaria (I?)
Trachemys stejnegeri stejnegeri (I) Graptemys pseudogeographica (W)
Anolis ferreus Pelusios castaneus (W)
Capitellum mariagalantae (PE) Terrapene carolina (W)
Gymnophthlamus underwoodi (I) Trachemys scripta elegans (I)
Hemidactylus mabouia (I?) Trachemys stejnegeri stejnegeri (W)
Iguana delicatissima (PX) Anolis roquet roquet
Iguana iguana (I) Anolis roquet caracoli
Sphaerodactylus fantasticus anidrotus Anolis roquet majolgris
Thecadactylus rapicauda Anolis roquet salinei
Alsophis antillensis (PX) Anolis roquet summus
Liophis juliae (PX) Anolis roquet zebrilus
Capitellum metallicum (PE)
DOMINICA BANK Gekko gecko (I)
Dominica (15.43°/-61.35°) (751 km2) Gonatodes albogularis (?)
Eleutherodactylus amplinympha Gymnophthalmus pleii
Eleutherodactylus johnstonei (I, E?) Gymnophthalmus underwoodi (I)
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 155

Hemidactylus mabouia (I?) Sphaerodactylus vincenti

Iguana delicatissima Thecadactylus rapicauda
Iguana iguana (I) Tetracheilostoma sp.
Leiocephalus herminieri (PE)
Îlet Chevalier (14.43°/-60.83°) (0.2 km2)
Mabuya mabouya (PE)
Eleutherodactylus johnstonei (I)
Sphaerodactylus vincenti festus
Anolis roquet saline
Sphaerodactylus vincenti josephinae
Gymnophthalmus pleii
Sphaerodactylus vincenti pheristus
Hemidactylus mabouia (I?)
Sphaerodactylus vincenti psammius
Sphaerodactylus vincenti psammius
Sphaerodactylus vincenti ronaldi
Bothrops lanceolatus (PX)
Sphaerodactylus vincenti ssp.
Thecadactylus rapicauda Îlet Frégate (14.61°/-60.86°) (0.01 km2)
Boa sp. (PE) Anolis roquet ssp.
Boa constrictor ssp. (W) Îlet Hardy (14.41°/-60.83)(0.026 km2)
Boa orophias (W) Anolis roquet salinei
Bothrops lanceolatus
Gymnophthalmus pleii
Bothrops caribbaeus (W)
Hemidactylus mabouia (I?)
Epicrates cenchria (W)
Liophis cursor (PE) Îlet Lavigne (= Gros Ilet) (14.63°/-60.89°) (0.05 km2)
Natrix natrix (W) Eleutherodactylus johnstonei (I)
Pantherophis alleghaniensis (W) Rhinella marina (I)
Pantherophis guttatus (W) Anolis roquet ssp.
Python regius (W) Gymnophthalmus pleii
Tetracheilostoma bilineatum Hemidactylus mabouia (I?)
Caiman sp. (W)
Îlet Long (14.61°/-60.86°) (0.12 km2)
Gros Îlet (14.55°/-61.01°) (0.12 km2) Eleutherodactylus johnstonei (I)
Eleutherodactylus johnstonei (I) Rhinella marina (I)
Anolis roquet ssp. Anolis roquet ssp.
Gymnophthalmus pleii Hemidactylus mabouia (I?)
Îlet à Eau (14.69°/-60.91°) (0.02 km2) Îlet Madame (14.67°/-60.88°) (0.03 km2)
Eleutherodactylus johnstonei (I) Anolis roquet ssp.
Anolis roquet ssp.
Îlet Métrente (14.62°/-60.86°) (0.06 km2)
Îlet Aigrettes (14.42°/-60.84°) (0.002 km2) Eleutherodactylus johnstonei (I)
Anolis roquet salinei Anolis roquet ssp.
Hemidactylus mabouia (I?) Gymnophthalmus pleii
Îlet aux Rats (14.68°/-60.90°) (0.005 km2) Hemidactylus mabouia (I?)
Anolis roquet ssp.
Îlet Oscar (14.63°/-60.85°) (0.04 km2)
Îlet Burgaux (14.41°/-60.84)(0.005 km ) 2
Eleutherodactylus johnstonei (I)
Hemidactylus mabouia (I?) Anolis roquet salinei x A. r. roquet
Îlet Cabrits (14.39°/-60.87°) (0.04 km2) Gymnophthalmus pleii
Anolis roquet salinei Hemidactylus mabouia (I?)
Petit Îlet (14.39°/-60.87°) (0.01 km2) Îlet Percé (14.41°/-60.84°) (0.005 km2)
Anolis roquet ssp. Anolis roquet salinei
Gymnophthalmus pleii Îlet Petite Grenade (14.57°/-60.84°) (0.05 km2)
Îlet Chancel (14.69°/-60.89°) (0.8 km ) 2
Anolis roquet ssp.
Eleutherodactylus johnstonei (I)
Îlet Petite Martinique (14.69°/-60.91°) (0.05 km2)
Eleutherodactylus martinicensis
Eleutherodactylus johnstonei (I)
Rhinella marina (I)
Anolis roquet ssp.
Anolis roquet roquet
Gymnophthalmus pleii Îlet Petit Vincent (14.69°/-60.92°) (0.005 km2)
Hemidactylus mabouia Eleutherodactylus johnstonei (I)
Iguana delicatissima Anolis roquet ssp.
156 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Îlet Poirier (14.41°/-60.84°) (0.021 km2) Boa orophias

Anolis roquet salinei Bothrops caribbaeus
Gymnophthalmus pleii Clelia errabunda (PE)
Hemidactylus mabouia (I?) Liophis ornatus (PX)
Tetracheilostoma breuili
Îlet Ragot ou Îlet de la Grotte (14.69°/-60.88)(0.03 km2)
Eleutherodactylus johnstonei (I) Dennery Island (13.91°/-60.88°) (0.02 km2)
Anolis roquet ssp. Anolis luciae
Hemidactylus mabouia (I?) Hemidactylus palaichthus
Îlet à Ramiers (14.54°/-61.08°) (0.02 km2) Maria Island (North) (13.73°/-60.93°) (0.02 km2)
Anolis roquet ssp. Cnemidophorus vanzoi
Sphaerodactylus vincenti ssp. Hemidactylus palaichthus
Iguana delicatissima (I) Sphaerodactylus vincenti
Thecadactylus rapicauda
Îlet Rat (14.68°/-60.90°) (0.01 km2)
Anolis roquet salinei Maria Island (South) (13.72°/-60.93°) (0.09 km2)
Anolis luciae
Îlet Saint-Aubin (14.77°/-60.96)(0.03 km2) Cnemidophorus vanzoi
Anolis roquet majolgris x A. r. roquet Gymnophthalmus pleii
Îlet Thierry (14.63°/-60.85°) (0.05 km2) Hemidactylus palaichthus (I)
Eleutherodactylus johnstonei (I) Sphaerodactylus vincenti
Anolis roquet ssp. Thecadactylus rapicauda
Gymnophthalmus pleii Liophis ornatus
Hemidactylus mabouia (I?) Tetracheilostoma breuili
Loup Garou (14.67°/-60.85°) (0.01 km2) Pigeon Island (14.09°/-60.96)
Anolis roquet ssp. Anolis luciae

Rocher du Diamant (14.44°/-61.04°) (0.06 km2) Praslin Island (13.88°/-60.89°) (0.01 km2)
Anolis roquet ssp. Eleutherodactylus johnstonei (I)
Gymnophthalmus pleii Anolis luciae
Sphaerodactylus vincenti adamas Cnemidophorus vanzoi (I)
Gymnophthalmus pleii
Thecadactylus rapicauda
Liophis cursor (PE) Rat Island (14.04°/-60.98)
Anolis luciae
ST. LUCIA BANK Scorpion Island (13.77°/-60.92°) (0.02 km2)
St. Lucia (13.92°/-60.96°) (616 km2) Anolis luciae
Eleutherodactylus johnstonei (I)
Eleutherodactylus martinicensis BARBADOS BANK
Leptodactylus fallax (PX) Barbados (13.14°/-59.56°) (430 km2)
Osteopilus septentrionalis (I) Eleutherodactylus johnstonei (I)
Rhinella marina (I) Rhinella marina (I)
Scinax ruber (I, E?) Chelonoidis carbonaria (I?)
Chelonoidis sp. (I?) Chelonoidis sp. (F)
Alinea luciae (PE) Trachemys scripta elegans (W)
Anolis extremus (I) Alinea lanceolata (PE)
Anolis luciae Ameiva ameiva (I)
Anolis wattsi wattsi (I) Anolis extremus
Gymnophthalmus pleii Anolis sagrei ssp. (I)
Hemidactylus mabouia (I?) Gymnophthalmus underwoodi (I)
Hemidactylus palaichthus Hemidactylus mabouia (I?)
Iguana iguana (I) Kentropyx borckiana
Iguana cf. iguana Phyllodactylus pulcher
Sphaerodactylus microlepis microlepis Liophis perfuscus (PE)
Sphaerodactylus vincenti diamesus Mastigodryas bruesi (I)
Thecadactylus rapicauda Ramphotyphlops braminus (I)
Amphisbaena fuliginosa (W?) Tetracheilostoma carlae
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 157

ST. VINCENT BANK Mastigodryas bruesi

St. Vincent (13.26°/-61.19°) (350 km )2 Tantilla melanocephala (I)
Eleutherodactylus johnstonei (I) Typhlops tasymicris
Pristimantis shrevei Crocodilus intermedius (W)
Leptodactylus validus (I) Baliceaux (12.94°/-61.14°) (1.0 km2)
Rhinella marina (I?) Chelonoidis carbonaria (I?)
Chelonoidis sp. (I?) Anolis aeneus
Ameiva ameiva Hemidactylus mabouia (I?)
Anolis griseus Iguana iguana
Anolis sagrei ssp. (I) Corallus grenadensis
Anolis trinitatus Mastigodryas bruesi
Copeoglossum aurae (PX)
Gymnophthalmus underwoodi (I) Baradal (12.63°/-61.35°) (0.02 km2)
Hemidactylus mabouia (I?) Anolis aeneus
Iguana iguana Battowia (12.96°/-61.13°) (0.7 km2)
Sphaerodactylus vincenti Anolis aeneus
Thecadactylus rapicauda Iguana iguana
Chironius vincenti Bequia (13.02°/-61.23) (18.0 km2)
Corallus cookii Eleutherodactylus johnstonei (I)
Mastigodryas bruesi Leptodactylus validus (I)
Chateaubelair (13.28°/-61.25°) (0.06 km2) Rhinella marina (I?)
Anolis trinitatis Chelonoidis carbonaria (I?)
Gymnopthalmus underwoodi (I) Ameiva ameiva tobagana
Iguana iguana Anolis aeneus
Anolis richardi
Milligan Cay (13.12°/-61.17°) (0.02 km2)
Bachia heteropus alleni
Iguana iguana
Copeoglossum aurae
Young Island (= Young’s Island) (13.12°/-61.20°) (0.04 km2) Gymnophthalmus underwoodi (I)
Anolis trinitatis Hemidactylus mabouia (I?)
Iguana iguana Iguana iguana
Marisora aurulae Sphaerodactylus kirbyi
Thecadactylus rapicauda
GRENADA BANK Corallus grenadensis
Grenada (12.12°/-61.68°) (311 km2) Mastigodryas bruesi
Eleutherodactylus johnstonei (I)
Canouan (12.71°/-61.32°) (7.4 km2)
Eleutherodactylus planirostris (W?)
Eleutherodactylus johnstonei (I)
Leptodactylus fallax (I, E)
Rhinella marina (I?)
Leptodactylus validus (I)
Chelonoidis carbonaria (I?)
Pristimantis euphronides
Ameiva ameiva tobagana
Rhinella marina (I?)
Anolis aeneus
Chelonoidis carbonaria (I?)
Anolis carolinensis or A. porcatus porcatus (W)
Ameiva ameiva tobagana
Anolis sagrei ssp. (I)
Anolis aeneus
Bachia heteropa alleni
Anolis richardii
Copeoglossum aurae or Marisora aurulae
Anolis sagrei ssp. (I)
Hemidactylus mabouia (I?)
Bachia heteropus alleni
Iguana iguana
Copeoglossum aurae (PX)
Corallus grenadensis
Gymnophthalmus underwoodi (I)
Mastigodryas bruesi
Hemidactylus mabouia (I?)
Iguana iguana Carriacou (12.48°/-61.45°) (32.0 km2)
Marisora aurulae (PX) Eleutherodactylus johnstonei (I)
Thecadactylus rapicauda Rhinella marina (I?)
Amphisbaena fuliginosa (W?) Chelonoidis carbonaria (I?)
Clelia clelia groomei Ameiva ameiva tobagana
Corallus grenadensis Anolis aeneus
158 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

Anolis richardii Mayreau (= Mayero) (12.64°/-61.39°) (2.6 km2)

Copeoglossum aurae Chelonoidis carbonaria (I?)
Hemidactylus mabouia (I?) Ameiva ameiva tobagana
Iguana iguana Anolis aeneus
Marisora aurulae Hemidactylus mabouia (I?)
Sphaerodactylus kirbyi Iguana iguana
Thecadactylus rapicauda Marisora aurulae
Corallus grenadensis Sphaerodactylus kirbyi
Mastigodryas bruesi Corallus grenadensis
Tantilla melanocephala (W) Mastigodryas bruesi
Caiman crocodilus (W)
Mustique (12.84°/-61.18°) (5.2 km2)
Catholic Island (12.66°/-61.40°) (0.06 km2) Eleutherodactylus johnstonei (I)
Anolis aeneus Osteopilus septentrionalis (I)
Iguana iguana Rhinella marina (I?)
Frigate Island (12.41°/-61.48°) (0.07 km2) Chelonoidis carbonaria (I?)
Ameiva ameiva tobagana Ameiva ameiva tobagana
Anolis aeneus Anolis aeneus
Bachia heteropa alleni
Glover Island (11.99°/-61.79°) (0.04 km2)
Copeoglossum aurae
Anolis aeneus
Gymnophthalmus underwoodi (I)
Marisora aurulae
Hemidactylus mabouia (I?)
Green Island (12.23°/-61.59°) (0.09 km2) Iguana iguana
Ameiva ameiva tobagana Sphaerodactylus kirbyi
Anolis aeneus Thecadactylus rapicauda
Thecadactylus rapicauda Corallus grenadensis
Hog Island (12.00°/-61.74°) (0.28 km2) Mastigodryas bruesi
Eleutherodactylus johnstonei (I) Ramphotyphlops braminus (W)
Anolis aeneus Tantilla melanocephala (W)
Gymnophthalmus underwoodi Palm Island (= Prune Island) (12.59°/-61.40°) (0.35 km2)
Hemidactylus mabouia (I?) Chelonoidis carbonaria (I?)
Isle à Caille (12.29°/-61.58°) (0.5 km2) Ameiva ameiva tobagana
Ameiva ameiva tobagana Anolis aeneus
Anolis aeneus Hemidactylus mabouia (I?)
Iguana iguana Iguana iguana
Isle à Quarte (12.96°/- 61.25°) (2.0 km2) Petit Canouan (12.78°/-61.28°) (0.05 km2)
Anolis aeneus Anolis aeneus
Iguana iguana Petit Dominique (12.51°/-61.40°) (0.3 km2)
Corallus grenadensis Anolis aeneus
Mastigodryas bruesi
Petit Martinique (12.52°/-61.38°) (0.7 km2)
Isle à Ronde (12.30°/-61.59°) (1.7 km2) Anolis aeneus
Ameiva ameiva tobagana
Copeoglossum aurae
Kick ‘em Jenny (= Diamond Island) (12.33°/-61.58°) (0.2 km2) Hemidactylus mabouia (I?)
Anolis aeneus Corallus grenadensis
Iguana iguana Mastigodryas bruesi
Large Island (12.41°/-61.49°) (0.5 km2) Petit Mustique (12.84°/-61.19°) (0.45 km2)
Anolis aeneus Ameiva ameiva ssp.
Les Tantes (12.32°/-61.55°) (0.18 km2) Anolis aeneus
Anolis aeneus Iguana iguana
Mabouya Island (12.49°/-61.49°) (0.06 km2) Petit Nevis (12.97°/-61.24°) (0.2 km2)
Anolis aeneus Hemidactylus mabouia (I?)
Anolis richardi Iguana iguana
Iguana iguana Sphaerodactylus kirbyi
POWELL AND HENDERSON: Island lists of West Indian amphibians and reptiles 159

Petite St. Vincent (12.54°/-61.38°) (0.35 km2)

Eleutherodactylus johnstonei (I)
Anolis aeneus
Hemidactylus mabouia (I?)
Iguana iguana
Corallus grenadensis
Mastigodryas bruesi
Ramphotyphlops braminus (W)
Sandy Island (12.22°/-61.59°) (0.07 km2)
Ameiva ameiva tobagana
Anolis aeneus
Savan Island (12.81°/-61.21°) (0.08 km2)
Anolis aeneus
Iguana iguana
Sugar Loaf (= Levera Island) (12.23°/-61.61°) (0.07 km2)
Anolis aeneus
Anolis richardi
The Sisters (12.30°/-61.60°) (0.02 km2)
Anolis aeneus
Tobago Cays
Jamesby (12.63°/-61.36°) (0.01 km2)
Anolis aeneus
Petit Bateau (12.63°/-61.36°) (0.05 km2)
Ameiva ameiva tobagana
Anolis aeneus
Marisora aurulae
Petit Rameau (12.64°/-61.36°) (0.06 km2)
Anolis aeneus
Union (12.60°/-61.44°) (8.1 km2)
Eleutherodactylus johnstonei (I)
Eleutherodactylus sp. (I)
Rhinella marina (W)
Chelonoidis carbonaria (I?)
Ameiva ameiva tobagana
Anolis aeneus
Bachia heteropa alleni
Copeoglossum aurae
Gonatodes daudini
Gymophthalmus underwoodi (I)
Hemidactylus mabouia (I?)
Iguana iguana
Sphaerodactylus kirbyi
Corallus grenadensis
Mastigodryas bruesi
Tantilla melanocephala (W)
Typhlops tasymicris
160 BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 51(2)

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from the northern Bahamas, with an osteological assessment of other Neotropical tortoises. 49(1):1–44. $8.00

Snelson Jr., F. F., T. J. Krabbenhoft, & J. M. Quattro. 2009. Elassoma gilberti, a new species of pygmy sunfish
(Elassomatidae) from Florida and Georgia. 48(4):119–144. $7.00

Zaspel, J. M., S. J. Weller, & R. T. Carde. 2008. A review of Virbia (formerly Holomelina) of North America.
48(3):59–118. $10.00

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