Introduction to Plant Physiology
● Plant physiology is a branch of plant science studying how plants live and function
● Stephen Hales is considered the father of plant physiology
● M
ain physiological processes: diffusion, imbibition, water & mineral absorption, transportation,
transpiration, photosynthesis, respiration, etc.
Essential Nutrients in Plants
● 17 inorganic elements are essential for plant growth and completion of life cycle
● Macronutrients (needed in larger amounts):
○ Carbon (C), Hydrogen (H), Oxygen (O) → non-mineral, from air and water
○ Nitrogen (N), Phosphorus (P), Potassium (K), Calcium (Ca), Magnesium (Mg), Sulfur (S)
● Micronutrients (needed in smaller amounts):
○ Iron (Fe), Manganese (Mn), Zinc (Zn), Copper (Cu), Molybdenum (Mo), Boron (B), Chlorine
(Cl), Sodium (Na)
● Some nutrients have special roles:
○ Silicon – grass plants
○ Cobalt – legumes
○ Sodium – C₄ plants
Mineral Absorption
● Minerals are absorbed in ionic form (e.g., K⁺, Ca²⁺, NO₃⁻, SO₄²⁻)
● Root hairs and meristematic regions are involved in absorption
● Two types of absorption mechani[Link]
○ Active absorption (requires ATP) – ions move against concentration gradient
○ Passive absorption – ions move along concentration gradient
Active Absorption Theories
● Carrier Concept (Vanden Honert, 1937):
○ Specific carrier molecules form complexes with ions and transport them
○ Uptake rate plateaus when all carriers are occupied
● Protein-Lecithin Theory (Bennet Clark, 1956):
○ Lecithin acts as carrier for ions
○ Enzyme lecithinase mediates the process
● Proton-Anion Co-transport Theory:
○ Proton motive force drives anion uptake along with H⁺ ions
○ Relies on pH and ATPase activity
● Cytochrome Pump Theory (H. Lund, 1954):
○ Involves dehydrogenase reactions and electron transport
○ Anions enter via redox activity of cytochromes
Passive Absorption Theories
● Mass flow theory – ions move with water due to transpiration
● Diffusion theory – ions diffuse from high to low concentration
● Ion Exchange theory – ions are exchanged between soil solution and root surfaces
○ E.g., H⁺/OH⁻ from roots displace cations/anions
● D
onnan Equilibrium – due to presence of non-diffusible ions, ions distribute unequally across
membranes
Factors Affecting Mineral Absorption
● Temperature – absorption is temperature-dependent
● Oxygen – essential for active absorption
● Light – enhances transpiration and indirectly affects absorption
● pH – acidic pH helps cation uptake; alkaline favors anions
● Presence of other ions – can inhibit or enhance uptake (ion antagonism)
● Growth activity – active cell division and elongation increase absorption