Butterflies of Western Ghats by Dr. Raju Kasambe FINAL 04 January 2016
Butterflies of Western Ghats by Dr. Raju Kasambe FINAL 04 January 2016
An e-Book
I
Butterflies of Western Ghats . 2016
Address:
B-205, Trimurti Apartments,
Borkar Lane, Lokmany Tilak Nagar,
Dombivli (East), Distt. Thane,
Maharashtra. PIN-421201.
Email: [email protected], [email protected]
Mobile: +91-9004924731.
Office/ Workplace:
Bombay Natural History Society,
Hornbill House, Dr. Salim Ali Chowk,
Shaheed Bhagat Singh Road,
Mumbai-400001, Maharashtra, INDIA
Off. Tel. 91-22-22821811 (Hornbill House).
Off. Tel. 91-22-22429477 (Conservation Education Center, Goregaon).
Recommended citation:
Kasambe, Raju (2016): Butterflies of Western Ghats. An e-Book. Self Published.
Pp.327+vii.
II
Butterflies of Western Ghats . 2016
Dedicated to
Two butterfly-passionate persons
and
III
Butterflies of Western Ghats . 2016
CONTENTS
Foreword………………………………………………………………………….V
Preface……………………………………………………………………………VI
Acknowledgements……………………………………………………………...VII
Hesperiidae—Skipper Butterflies…………………………………………..19–72
Papilionidae—Swallowtail Butterflies………………………………….......73–92
Lycaenidae—Blues………………………………………………………..125–210
Nymphalidae—Brush-footed Butterflies………………………………...211–300
References……………………………………………………………………….326
IV
Butterflies of Western Ghats . 2016
Foreword
I think, this is the first of its kind e-Book on the subject, as books
which are accessible to the masses are need of the day. Now a day,
even Android Apps are becoming an important source of information.
The e-Book has an entire chapter highlighting the importance of the
Western Ghats as a ‘Biodiversity Hotspot’. The diversity and
endemicity of butterflies found here drives home the point that the
entire Western Ghats should be conserved at any cost.
I am sure the e-Book will be useful to all, including butterfly lovers and
naturalists.
Asad R. Rahmani
V
Butterflies of Western Ghats . 2016
Preface
This e-Book is being published solely for the educational purpose and no commercial
gains are expected out of it. The e-Book is free for downloading and distribution and
does not bear any price. I urge nature lovers and butterfly photographers to help with
better quality photographs of the remaining species of butterflies found in the Western
Ghats, so that the second edition of this e-Book will be more comprehensive and better.
Photographs taken by me are copyrighted under the Creative Commons Attribution 3.0
License. These can be used by anybody for educational purpose. However, for
photographs taken by others, the copyright remains with them. I suggest that the
individual photographers be contacted, or the appropriate licenses be checked before
they are used by anybody.
I request experts to suggest improvements in the e-Book. Please inform me if you find
any mistakes in the e-Book, which can be rectified in the second edition.
I hope this e-Book will be of use to the young nature lovers, it will give them company at
the most inaccessible places. I look forward to your help by providing photographs of
missing species and suggestions for the second edition of this e-Book.
Raju Kasambe
VI
Butterflies of Western Ghats . 2016
Acknowledgements
My sincere thanks to the following friends (in alphabetical order) who have contributed their
beautiful and photographs of rare butterfly species for this e-Book. Some of the friends literally
opened up their vast collections for me to take any photographs. I am grateful to all of them.
Without photographs the e-Book would not have been useful. Many of them have selflessly
contributed their innumerable photographs to Wikipedia and Wikimedia Commons making
them available to the world without any expectations or gains.
Aditya Joshi, Dr. Amol Patwardhan, Anila Manalil, Aparajita Datta, Ashwatha K.N., Ashok
Sengupta, Ayan Chakraborty, Blaise Pareira, Chinmayi S.K., Dattaprasad Sawant, David Raju,
Delip Kumar Das, Dhaval Momaya, Divakar Thombre, Dr. Amol Patwardhan, Dr. Jayant
Wadatkar, Dr. M.S. Mayilavahanan, Dr. Tarique Sani, Dr. V.C. Balakrishnan, Erin Silversmith,
Gaurab Nandi Das, Gopakumar, Harishchandra Mhatre, Hariharan Subramanian, Hemant Ogale,
J.M. Garg, Kalyan Varma, Jeevan Jose, K. Mohan Raj, Kishen Das, L. Shyamal, Makarand
Kulkarni, Mandar Sawant, Manidip Mandal, Manu Gangadhar, Marvelyn Dias, Morry G., Mukul
Hinge, Nandish Songire, Parag Giri, Parag Rangnekar, Parixit Kafley, Pinakin Karve, PKG Mohan,
Pranav Gokhale, Pranav Ovalekar, Prashanth Bhat, Praveen G. Nair, Praveen J., R. Ravi, Raghu,
Rahul K. Natu, Rajendran TM, Rajkamal Goswami, Ravi Vaidyanathan, Richard Parker, Rohith
Sanjay, Sagar Sarang, Saish Borkar, Sanket Mhatre, Shyam Ghate, Sneha, Shreesh Deshpande,
Subhajit Mazumder, Subhiksha S., Sujit Borkar, Tarun Karmakar, Uajith, Varun Omanakuttan,
Vedant Kasambe, Vengolis, Vijay Barve, Vinayraj, Vivek Puliyeri, Yathin S. Krishnappa and
Yuwaraj Gurjar.
I want to thank the two websites (www.ifoundbutterflies.org and www.flutters.org) for making
an exhaustive knowledge bank on Indian butterflies available to all.
Dr. Deepak Apte (Director, Bombay Natural History Society) has always inspired me to write
books against all odds. I learnt a lot from Dr. Asad Rahmani (Senior Scientific Advisor, BNHS)
and I feel indebted to him for his guidance. Guidance from Mr. Isaac Kehimkar (butterfly expert)
about butterflies has immensely helped me in working on this e-Book. Isaac’s books had always
been the most referred and inspiring source for me.
I had written most part of this e-Book in 2012, when I published my Marathi book on
‘Maharashtratil Phulpakhare’ (aka Butterflies of Maharashtra), but the book got delayed due to
other priority work at hand. Support by many friends and colleagues at BNHS and my family has
always helped me in keeping myself motivated.
Raju Kasambe
VII
Butterflies of Western Ghats: Dr. Raju Kasambe Introduction 2016
Butterflies are classified under the Order Lepidoptera, together with the moths.
Egg
Like humans, there are male butterflies and female butterflies. After the
fertilization of the female’s eggs by the
male’s sperm, the female deposits the eggs
on leaves or stems of plants. This is the first
stage in the life cycle of a butterfly - the egg
stage. Butterflies have their own choice of
plant on which to lay eggs, they may lay eggs
on one or many species of plants. But, they
do not lay eggs on every plant. The eggs of
Egg of Common Palmfly by Raju Kasambe each species are different in shape and size
and can be spherical or oval etc. This stage
lasts from days to weeks.
Caterpillar or Larva
The second stage occurs when then the egg hatches and gives rise to the butterfly
larva or caterpillar. At this stage, which may
last for several days or weeks, the larva
spends its time eating and sleeping. Most
butterfly caterpillars eat plant leaves but a
few caterpillars can eat insects (e.g. Apefly
in India).
1
Butterflies of Western Ghats: Dr. Raju Kasambe Introduction 2016
instar signifies moulting of the skin (cuticle) by the caterpillar. Instars are
important because they allow the caterpillar to grow. The butterfly undergoes
about 4 to 6 moultings before finally going to the next stage, i.e. pupation.
Chrysalis or Cocoon
The third stage involves the transformation of the larva into chrysalis or pupa.
When the larva has grown to its full size,
it seeks a safe location for pupation. This
location is usually the underside of the
leaf but can also occur in the stems or
secluded place on a tree trunk.
Pupation may last for 2–3 weeks, after which the butterfly adult emerges from the
cocoon.
Butterfly
The fourth and final stage is the adult stage. It is a delight to see an adult butterfly
emerge from the cocoon. However, the
butterfly cannot fly immediately after
emerging out of the cocoon. It spends
the sometime (few hours or minutes)
drying its body and wings before it
finally takes off.
2
Butterflies of Western Ghats: Dr. Raju Kasambe Introduction 2016
forewings and another pair of hindwings. When the butterfly is already an adult, it
can gather nectar from flowers for its food and reproduce. It can give rise to other
beautiful butterflies.
Morphology of a Butterfly
Here are the meaning of the terminology used to describe various body parts of a
butterfly.
Head
Anterior portion of the butterfly’s body containing the sensory organs and the
brain.
Thorax
Portion of the butterfly’s body divided into three segments; it contains the motor
appendages, such as the legs and wings.
3
Butterflies of Western Ghats: Dr. Raju Kasambe Introduction 2016
Wing vein
Protruding line that gives the wing its rigidity and enables the blood to circulate.
Cell
Constituent element of a butterfly’s wing contained between the wing veins.
Antenna
Sensory organ made up of several segments and having mainly tactile and
olfactory functions.
Compound eye
Organ of vision made up of thousands of facets that perceive shapes, colors,
motion and distance.
Proboscis
Mouthlike part allowing the butterfly to feed through aspiration; the proboscis
folds back onto itself to avoid interfering with flight.
Foreleg
Articulated member attached to the first segment of the thorax and having
powerful sensory organs.
Middle leg
Large articulated member attached to the central segment of the thorax and having
powerful sensory organs.
Hind leg
Large articulated member attached to the terminal segment of the thorax and
having powerful sensory organs.
Abdomen
Posterior portion of the butterfly’s body made up of 10 segments and containing
the major vital organs, such as the heart, the intestines and the genital organs.
Hind wing
Appendage of flight attached to the terminal segment of the thorax.
4
Butterflies of Western Ghats: Dr. Raju Kasambe Introduction 2016
Forewing
Appendage of flight attached to the central segment of the thorax.
Parts of wings
Here is an illustration showing parts of butterfly wings.
5
Butterflies of Western Ghats: Dr. Raju Kasambe Introduction 2016
Butterflies are diurnal (active during the day) whereas moths are nocturnal. There
are some exceptions to this, as some butterflies are crepuscular and some moths are
diurnal.
Butterfly antennae are thin with club-shaped tips, compared with the feathery or
comb-like antennae of moths.
While at rest, butterflies usually fold their wings back over the bodies, while moths
flatten their wings against their bodies or spread them out in a "jet plane" position.
Butterflies form chrysalises (or cocoons) which are hard smooth and silkless.
Moths make cocoons that are wrapped in silk coverings.
6
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats - Butterflies 2016
Evergreen forests in Western Ghats in Periyar Tiger Reserve in Kerala by Manu Gangadhar.
Based on the distribution and status of butterflies and their host plants, Western
Ghats can be divided into three biogeographical parts (Gaonkar 1996).
Southern Western Ghats: The southern and most important part starts from just
north of Nagacoil (in Tamil Nadu) up to the Palghat gap. This area is home to the
highest number of species as well as endemics. Unfortunately, the lowland
evergreen forests in this area have mostly been replaced with plantations other
anthropogenic pressures exist here. The unique species of this area are Red-disc
Bushbrown Mycalesis oculus and Palni Fourring Ypthima ypthimoides. This area is
home to around 317 species of butterflies.
Centrel Western Ghats: The second important part starts north of the Palghat
Gap from the Nilgiri Wynaad area to South Goa. However, some endemic species
7
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats - Butterflies 2016
Northern Western Ghats: The third part is in Maharashtra and south Gujarat.
This area is has less diversity with around 200 species reported north of Amboli
Ghat. Many species from the central Western Ghats are not found here.
Karnataka, Goa and Maharashtra will also increase their species numbers once
study becomes more intensive and systemic.
8
Butterflies of Western Ghats: Dr. Raju Kasambe Importance of Butterflies 2016
Butterflies may be better indicators of the health of our environment than birds.
— Roger Tory Peterson
Intrinsic value
Butterflies and moths are intrinsically valuable and are worthy of conservation in their own right.
Butterflies and moths are part of the life on the earth and an important component of its rich
biodiversity. They have been around for at least 50 million years and probably first evolved some
150 million years ago. Butterflies and moths are a highly diverse group comprising over 250,000
species and make up around one quarter of all named species.
Aesthetic value
Butterflies and moths are beautiful. We humans love all beautiful things in nature. Butterflies
add to the aesthetic value of any natural landscape and heritage. Many butterflies are iconic and
popular. Butterflies have been studied for over 300 years by humans.
9
Butterflies of Western Ghats: Dr. Raju Kasambe Importance of Butterflies 2016
Educational value
Butterflies and moths have fascinating life-cycles that are used in many countries to teach
children about the natural world. The transformation from egg to caterpillar to chrysalis is one of
the wonders of nature.
Other educational aspects include the intricate wing patterns and iridescence, and as examples of
insect migration.
Scientific value
Butterflies (and moths to a lesser extent) are an extremely important group of ‘model’ organisms
used, for centuries, to investigate many areas of biological research, including such diverse fields
as navigation, pest control, embryology, mimicry, evolution, genetics, population dynamics and
biodiversity conservation.
The long history and popularity of butterfly study have provided a unique data resource on an
insect group unmatched in geographical scale and timescale anywhere in the world. This has
proved extremely important for scientific research on climate change.
Ecosystem value
Butterflies and moths are indicators of a healthy environment and healthy ecosystems. They
indicate a wide range of other invertebrates, which comprise over two-thirds of all species. Areas
rich in butterflies and moths are rich in other invertebrates. These collectively provide a wide
range of environmental benefits, including pollination and natural pest control.
Moths and butterflies are an important element of the food chain and are prey for birds (like bee-
eaters, drongos and flycatchers), bats and other insectivorous animals (like lizards and geckos).
Butterflies and moths support a range of other predators and parasites (like the parasitoid wasps),
many of which are specific to individual species, or groups of species.
Health value
People enjoy seeing butterflies both around their homes and in the countryside. Social media
groups of butterfly lovers in India have more than 20,000 members. These people photograph,
survey and share butterfly information on social media. They travel thousands of kilometers in
search of butterflies. Now, butterfly gardening is becoming an important passion and many
butterfly gardens are in the making in India. People feel distressed in the company of butterflies
and they love to watch and photograph them.
Economic value
10
Butterflies of Western Ghats: Dr. Raju Kasambe Importance of Butterflies 2016
Thousands of people visit India each year looking for butterflies. Eco-tourism and now butterfly
tourism is bringing in valuable income to many local tribes and butterfly garden owners across
the country. Sikkim, Arunachal Pradesh, Assam, Kerala and Maharashtra have become hubs of
butterfly tourism and related businesses.
Every butterfly and moth has developed its own suite of chemicals to deter predators and
parasites, find a mate, and overcome the chemical defences of its host plant. Each of these
chemicals has a potential value and could be exploited economically.
Conservation of butterflies
In India, study and research on butterflies is still at the basic level. We are still working mostly
on the diversity inventories of various places. There are some good research papers coming out
on other aspects of Lepidoptera.
We have few names for butterflies in regional languages, but people do not have detailed
information about butterflies. People’s awareness regarding butterflies and their conservation is
almost next to nothing. It is for this reason butterflies are given little importance in the issues of
wildlife conservation.
Only recently, in June 2015, Maharashtra declared Blue Mormon Papilio polymnestor as the
‘State Butterfly’. Recently, in August 2016, Karnataka also declared Southern Birdwing Troides
minos as its ‘State Butterfly’ recognizing the importance of the butterflies in nature.
Now a good amount of academic research on butterflies is taking place in India but we do not
have good bibliographies on published information or baseline data on butterflies. For many of
our protected areas we have inventories for several other biological groups (mammals, birds,
reptiles, trees etc.), most of our parks and sanctuaries do not even have butterfly lists.
Moreover, very few serious ecological studies on the Lepidoptera of the region have been
undertaken and thus very little technical information is available for managers and policy makers
to take steps for effective butterfly conservation.
Habitats have been and are being destroyed on a massive scale. Now with the looming threat of
climate change and increasing pollution of the atmosphere may result into the disappearance or
drastic decline in the numbers of butterflies in many areas.
Conserving butterflies will improve our whole environment for wildlife and enrich the lives of
people now and in the future.
11
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats 2016
12
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats 2016
Western Ghats are home to many hill stations like Matheran, Lonavala-Khandala,
Mahabaleshwar, Panchgani, Amboli Ghat, Kudremukh and Kodagu. The extreme
northern parts of Western Ghats falls in the Dangs district of Gujarat, known for
Dang (Bamboo) forests. The confluence of the Eastern and the Western Ghats is at
Biligirirangan Hills in Karnataka. Anamudi at 2,695 metres in Kerala is the highest
peak in Western Ghats. Mullayanagiri is the highest peak in Karnataka at 1,950
meters.
The smaller ranges of the Western Ghats include the Cardamom Hills and the
Nilgiri Hills. Cardamom hills are located in southeast Kerala and southwest Tamil
Nadu. They conjoin the Anaimalai Hills to the northwest, the Palni Hills to the
northeast and the Agasthyamalai Hills to the south as far as the Ariankavu pass.
The crest of the hills forms the boundary between Kerala and Tamil Nadu.
Anamudi is also located in Cardamom Hills. The Nilgiri Hills are home to the hill
station Ooty. There are many important passes in Western Ghats such as Tamhini
Ghat, Palakkad Gap, Naneghat, Kasara ghat etc.
The northern portion of the narrow coastal plain between the Western Ghats and
the Arabian Sea is known as the Konkan Coast, the central portion is called Kanara
and the southern portion is called Malabar region or the Malabar Coast. The
foothill region east of the Ghats in Maharashtra is known as Desh, while the
eastern foothills of central Karnataka state is known as Malenadu.
mammal species, 15 bird species (now 26), 43 amphibian species, 5 reptile species
and 1 fish species. Of the total 325 globally threatened species in the Western
Ghats, 129 are classified as Vulnerable, 145 as Endangered and 51 as Critically
Endangered (Source: http://whc.unesco.org/en/list/1342).
Rivers
The rivers that originate in Western Ghats and flow towards west are Periyar,
Bharathappuzha, Netravati, Sharavathi, Mandovi etc. The west flowing rivers of
Western Ghats are fast-moving, owing to the short distance travelled and steeper
gradient. The steep gradient makes the Jog Falls on Shravasthi River in Karnataka
as one of the most spectacular waterfalls in India. The rivers that originate in
Western Ghats and flow towards east include three major rivers viz. Godavari,
Krishna and Kaveri, and many smaller tributary rivers such as Tunga, Bhadra,
Bhima, Malaprabha, Ghataprabha, Hemavathi, Kabini. These east flowing rivers
are comparatively slower moving and eventually merge into larger rivers such as
the Kaveri and Krishna.
Climate
Climate In comparison to the eastern side, the western side of the Western Ghats is
area of high rainfall because the mountains intercept the rain-bearing westerly
monsoon winds. The dense montane forests also contribute to high precipitation.
The climate is humid and tropical in the lower reaches tempered by the proximity
to the sea. Elevations of 1,500 m and above in the north and 2,000 m and above in
the south have a more temperate climate.
Average annual temperature here is around 15 °C. In some parts frost is common,
and temperatures touch the freezing point during the winter months. Mean
temperature range from 20 °C in the south to 24 °C in the north. It has also been
observed that the coldest periods in the south Western Ghats coincide with the
wettest. During the monsoon season between June and September, the unbroken
Western Ghats chain acts as a barrier to the moisture laden clouds. The heavy,
eastward-moving rain-bearing clouds are forced to rise and in the process deposit
most of their rain on the windward side. Rainfall in this region averages 3,000–
4,000 mm. The eastern region of the Western Ghats which lie in the rain shadow,
receive far less rainfall averaging about 1,000 mm bringing the average rainfall
figure to 2,500 mm.
14
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats 2016
Vegetation
Due to a sharp contrast in precipitation between western and eastern slopes of the
Western Ghats, there is a clear difference between the vegetation of the two sides.
Similarly, there is also a clear contrast between the northern and southern Western
Ghats. Moreover, the vegetation found on the high hills is also different from the
low hills. Thus, there are various kinds of vegetations found in Western Ghats as
follows: The western slopes have tropical and subtropical moist broadleaf forests
marked predominantly by Rosewood, Mahogany, Cedar etc. These slopes appear
green in almost all parts of the year. No time is fixed when these trees would shade
their leaves. The eastern slopes of the Western Ghats have dry as well as moist
deciduous forests marked predominantly by Teak, Sal, Shisham, Sandalwood etc.
trees. Further, on the northern side of the Wayanad forests; we find dry deciduous
forests while on the southern side there are wet deciduous forests. The evergreen
Wayanad forests of Kerala mark the transition zone between the northern and
southern ecoregions of the Western Ghats.
The southern ecoregions are generally wetter and more species-rich. South
Western Ghats Montane rain forests are the most species-rich ecoregions in
peninsular India. Eighty percent of the flowering plant species of the entire
Western Ghats range are found in this ecoregion. The areas which are high in
elevation are cooler and wetter in the north and so the forests there are called North
Western Ghats Montane rain forests. The vegetation here is evergreen
characterized by trees of family Lauraceae. Such plants include Litsea glutinosa or
Maida lakri in Hindi (a plant of medicinal value), Cinnamomum (Tejpatta) etc.
There are montane grasslands as well as stunted forests also in the Western Ghats.
The forest in the Western Ghats has been severely affected due to human activities,
especially clear felling for tea, coffee, and teak plantations during 1860 to 1950.
Species that are rare, endemic and habitat specialists are more adversely affected
and tend to be lost faster than other species. Complex and species rich habitats like
the tropical rainforest are much more adversely affected than other habitats. The
area is ecologically sensitive to development. Though this area covers barely five
percent of India’s land, 27% of all species of higher plants in India (4,000 of
15,000 species) are found here. Almost 1,800 of these are endemic to the region.
15
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats 2016
Protected Areas
Fauna
There are two biodiversity hotspots in our country viz. Eastern Himalayas and
Western Ghats. Western Ghats are home to over 5000 species of flowering plants,
139 mammal species, 508 bird species and 179 amphibian species, many
undiscovered species lives.
Mammals
There are at least 139 mammal species. Of the 16 endemic mammals found in
Western Ghats, 13 are threatened. Following four are important species endemic to
Western Ghats. Malabar Large-spotted Civet or just Malabar Civet Viverra
civettina (Critically Endangered), Lion-tailed Macaque Macaca silenus
(Endangered), Brown Palm Civet Paradoxurus jerdoni (Least Concerned) and
Nilgiri Tahr Nilgiritragus hylocrius (Endangered).
Birds
There are more than 500 bird species reported from Western Ghats. There are at
least 26 species of birds endemic to the Western Ghats including the Nilgiri
Imperial-pigeon Ducula cuprea, Nilgiri Woodpigeon Columba elphinstonii, Grey-
fronted Green-pigeon Treron affinis, Malabar Parakeet (Blue-winged Parakeet)
Psittacula columboides, Malabar Grey Hornbill Ocyceros griseus, Malabar Barbet
(Crimson-fronted Barbet) Xantholaema malabarica, Nilgiri Pipit Anthus
nilghiriensis, Malabar Woodshrike Tephrodornis sylvicola, Flame-throated Bulbul
(Ruby-throated Yellow Bulbul) Pycnonotus gularis (Rubigula gularis), Grey-
headed Bulbul Pycnonotus priocephalus, Nilgiri Thrush Zoothera neilgherriensis,
16
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats 2016
Amphibians
Western Ghats is home to at least 179 amphibian species as many more are being
described and more than 80% of these are endemic to the rainforests of the
mountains. The region is also home to many endemic caecilian species.
Fish, Insects
As of 2004, 288 freshwater fish species are listed from the Western Ghats,
including 35 which are also known from brackish or marine water. Several new
species have been described from the region since then.
Insects
Western Ghats is home to roughly 6,000 insect species. This includes 334 species
of butterflies. The Western Ghats is home to 174 species of odonates (107
dragonflies and 67 damselflies), including 69 endemics. Most of the endemic
odonates are closely associated with rivers and streams, while the non-endemics
typically are generalists. There are several species of leeches found all along the
Western Ghats.
Flora
Of the 7,402 species of flowering plants occurring in the Western Ghats, 5,588
species are native or indigenous and 376 are exotics naturalised and 1,438 species
are cultivated or planted as ornamentals. Among the indigenous species, 2,253
species are endemic to India and of them, 1,273 species are exclusively confined to
the Western Ghats.
17
Butterflies of Western Ghats: Dr. Raju Kasambe Western Ghats 2016
Sources:
http://www.gktoday.in/western-ghats/
http://whc.unesco.org/en/list/1342
https://en.wikipedia.org/wiki/Western_Ghats
Map: https://i2.wp.com/www.insightsonindia.com/wp-
content/uploads/2012/07/western-ghats-detailed-map.jpg
18
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Family Facts
Hesperiidae: The Skippers
Butterflies belonging to the family Hesperiidae are known as the “skippers”
because the butterflies exhibit a rapid, erratic or "skipping" flight pattern. The
Skippers are not considered to be "true" butterflies, but are more closely related to
the true butterflies than are the moths.
Many species of Skippers has a habit of basking in the sunlight with a unique
posture, the forewings being open only half way and the hindwings open fully.
This gives them a "fighter-jet" like appearance. They are largely brown, orange and
tawny.
Many species have very long proboscis which makes it possible for them to get
nectar from flowers with long tubular corolla. Many fly at dawn or dusk but others
fly during the daytime.
Eggs are tiny (less than the width of the head of a pin!) and vary in appearance, but
often are dome-shaped. Caterpillars generally are green with tapered bodies, often
live in shelters constructed with leaves and silk. Pupae hang in loose cocoons and
may have a silk girdle.
19
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution:
Throughout India.
20
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
B. j. fergusonii de Niceville, [1894]: Maharashtra to Kerala.
21
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Anila Manalil
Distribution: Subspecies:
B. g. kanara (Evans, 1926): Goa to Kerala.
22
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
B. s. sena (Moore, [1866]): Maharashtra to Madhya Pradesh and south to Kerala,
Himachal Pradesh to N.E. India; Andaman & Nicobar Is.
23
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
C. b. benjaminii (Guerin-Meneville, 1843): Karnataka to Kerala.
24
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
H. b. badra (Moore, [1858]): Maharashtra to Kerala; North Bihar to N.E. India;
Andaman & Nicobar Is.
25
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution:
H. c. chromus (Cramer, [1780]): Throughout India and Andaman & Nicobar Is.
26
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Tarun Karmakar
Distribution: Subspecies:
H. t. taminatus (Huebner, 1818): Karnataka to Kerala.
27
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Dattaprasad Sawant
Distribution: Subspecies:
H. v. indica Evans, 1932: Sikkim to N.E. India; Goa to Kerala.
28
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Anila Manalil
Distribution:
Maharashtra to Kerala; N.E. India.
29
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
A. d. dioscorides (Fabricius, 1793): Maharashtra east to West Bengal and south to
Kerala; Himachal Pradesh to N.E. India.
30
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
A. v. nilgiriana (Moore, 1883): Tamil Nadu to Kerala.
Subspecies:
A. v. vindhiana (Moore, 1883): Rajasthan to Tamil Nadu; Madhya Pradesh.
31
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
B. f. farri (Moore, 1878): Maharashtra southwards to Kerala and eastwards to W.
Bengal; Uttarakhand to N.E. India.
32
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Gujarat east to W. Bengal and southwards to Kerala; Jammu & Kashmir to N.E.
India.
33
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Throughout India except Jammu & Kashmir.
34
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Maharashtra to Kerala; Jharkhand; Sikkim to N.E. India.
35
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
H. h. hindu Evans, 1937: Karnataka; Kerala; Tamil Nadu.
36
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Maharashtra to West Bengal and southward to Kerala; Uttarakhand to N.E. India;
Andaman & Nicobar Is.
37
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
I. s. luteipalpis (Ploetz, 1886). Gujarat to Kerala.
38
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution:
Gujarat to West Bengal and southward to Kerala; Delhi; Uttarakhand to N.E India.
39
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution:
G. t. thyrsis (Fabricius, 1775): Maharashtra to Kerala; Andhra Pradesh; to N.E.
India, Himachal Pradesh; Andaman & Nicobar Is.
40
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
N. c. curvifascia (C. & R. Felder, 1862): Andaman & Nicobar Is. (Andamans);
Maharashtra to Kerala; Uttarakhand to N.E. India.
41
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Jeevan Jose
Distribution: Subspecies:
N. p. mangla Evans, 1939: Maharashtra to Kerala; Andhra Pradesh.
42
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
P. c . narooa (Moore,1878): Gujarat to Kerala and Jharkhand.
43
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Larval Host Plants: Oryza sativa (Rice), Cymbopogon nardus (Citronella Grass),
Imperata cylindrica (Blady Grass), Saccharum officinarum (Sugarcane).
Distribution:
Throughout India including Andaman & Nicobar Islands (Central Nicobars).
44
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
P. f. subfasciatus (Moore, 1878): Karnataka to Kerala; N.E. India.
45
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
S. g. gremius (Fabricius, 1798): Throughout India.
46
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
T. m. sagara (Moore, [1866]): Gujarat to Kerala; Madhya Pradesh;W. Bengal;
Jammu & Kashmir to N.E. India.
47
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Subspecies: T. c. ceramas (Hewitson, 1868): Karnataka, Kerala and Andhra
Pradesh.
Subspecies: T. c. media Evans, 1934: Goa and Karnataka.
Subspecies: T. c. nicevillei Watson, 1893: Maharashtra.
Subspecies: T. c. oberthueri Elwes & Edwards, 1897: Tamil Nadu (Anaimalai
Hills).
48
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Throughout India except Rajasthan.
49
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
T. c. colon (Fabricius, 1775): Gujarat to W. Bengal and southwards to Kerala;
Delhi to Uttar Pradesh; Uttarakhand to Sikkim.
50
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
C. c. oceanica (Mabille, 1904): Andaman & Nicobar Is. (Andamans), Sikkim to
N.E. India; west coast of India.
51
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Goa to Kerala.
Endemicity:
Endemic to Western Ghats.
52
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Gujarat to West Bengal and south to Kerala; Himachal Pradesh to N. E. India.
53
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Chinmayi S.K.
Distribution:
H. a. praba (Moore, [1866]): Himachal Pradesh to N. E. India; Andaman Is.; Goa
to Kerala.
54
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Maharashtra to Kerala; Sikkim to N.E. India.
55
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Clockwise from left: Male UN, DSF and WSF by Raju Kasambe
Distribution:
C. r. potiphera Hewitson, 1873: Gujarat east to Jharkhand and South to Kerala.
56
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN and UP by Gopakumar
Distribution:
C. a. agama (Moore, [1858]): Maharashtra east to Jharkhand and southward to
Kerala; Uttarakhand to N.E. India.
57
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN and UP by Gopakumar
Distribution: Subspecies:
C. a. vespa Evans, 1949: Tamil Nadu to Madhya Pradesh and Odisha.
58
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Raju Kasambe
Distribution:
Peninsular India, south from Gujarat and Madhya Pradesh.
59
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Raju Kasambe
Distribution:
Gujarat eastwards to West Bengal and southwards Kerala; Jammu and Kashmir to
N.E India; Andaman & Nicobar Is.
60
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
P. d. dan (Fabricius, 1787): Gujarat to Kerala; Andhra Pradesh.
61
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
C. i. indra Evans, 1926: Gujarat eastwards to West Bengal and southward to
Kerala.
62
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Tarun Karmakar
Distribution: Subspecies:
G. b. bhagava (Moore, [1866]): Goa to Jharkhand and south to Kerala; Sikkim to
N.E. India.
63
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
G. e. albofasciata Moore, 1879: Himachal Pradesh; Maharashtra to Kerala and
Telangana.
64
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Dhaval Momaya
Distribution:
O. a. angulatum (C. & R. Felder, 1862): Maharashtra to Kerala; Himachal Pradesh
to N.E. India.
65
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Raju Kasambe
Distribution: Subspecies:
S. d. adona Evans, 1949: Rajasthan to Odisha and south to Andhra Pradesh.
Subspecies:
S. d. davidsoni Swinhoe, 1912: Gujarat to Kerala.
66
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Vinayraj
Distribution: Subspecies:
S. p. hopkinsi Evans, 1921: Karnataka; Tamil Nadu.
Subspecies:
S. p. pandra Evans, 1949: Kerala to Rajasthan.
67
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution:
Throughout India.
68
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Anila Manalil
Distribution: Subspecies:
T. g. silvia Evans, 1934: Maharashtra to Kerala. Chhota Nagpur?
69
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Ayan Chakraborty
Distribution:
Subspecies: T. j. obscurus Mabille, 1876: Gujarat to Kerala.
70
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
UP by Raju Kasambe
Distribution: Subspecies:
T. l. litigiosa Fruhstorfer, 1910: Himachal Pradesh to N.E. India; Maharashtra to
Andhra Pradesh and Kerala; Jharkhand.
71
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Hesperiidae 2016
Distribution: Subspecies:
T. t. thwaitesi Moore, [1881]: Gujarat to Kerala and Andhra Pradesh.
72
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Family Facts
Family Papilionidae: Swallowtails
The Swallowtails are large butterflies having tails at the rear of the hindwing.
They are often black and yellow in color.
Swallowtails are generally characterized by the following: 1) have tails at the rear
of the hindwing; 2) medium to large in size; 3) unique pattern of wing veination on
the wings; and 3) fully developed and functioning forelegs in both sexes.
Eggs of Swallowtails are round and green. The young caterpillars of Swallowtails
often look like a bird dropping while older caterpillars are often greenish and
marked with large eyespots. They are equipped with an osmeterium, a Y-shaped
gland located behind the head which pops out and emit a pungent chemical to ward
off potential predators. Swallowtail pupae often have both a cremaster and a silk
girdle. Most beautiful (like Peacocks), threatened (like Kaiser-i-Hind, Bhutan
Glory) and largest of the butterflies (like the Birdwings) belong to this family.
These are much coveted as trophies and hence are illegally traded for their beauty.
73
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
74
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution: Subspecies:
G. d. eleius (Fruhstorfer, 1907): S. India to W. Bengal.
75
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution: Subspecies:
G. a. menides (Fruhstorfer, 1904): Kerala to Gujarat and W. Bengal.
76
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
UN by Anila Manalil
Distribution: Subspecies:
G. a. alcibiades (Fabricius, 1787): Western Ghats as far north as Goa.
77
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
G. n. nomius (Esper, 1799): Delhi; Rajasthan; Sikkim; Uttarakhand; Uttar Pradesh;
Bihar; throughout drier parts of Southern India to W. Bengal.
78
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution: Subspecies:
P. c. clytia Linnaeus, 1758: Throughout India except Jammu & Kashmir, Punjab
and Rajasthan, below 2750 m elevation.
79
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Female form stichius by Jeevan Jose Female form stichius by Raju Kasambe
Distribution: Subspecies:
P. p. romulus Cramer, [1775]: Throughout India below 2000 m elevation.
80
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution:
Western Ghats as far north as Goa.
Endemicity:
Endemic to Western Ghats.
81
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution: Subspecies:
P. h. daksha Hampson, 1889: Western Ghats south of Gujarat.
82
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Endemicity:
Endemic to India and Sri Lanka.
Distribution:
P. p. polymnestor Cramer, [1775]: Peninsular India as far north asW. Bengal and
Bangladesh, to Madhya Pradesh and Gujarat.
83
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution:
Western Ghats as far north as Goa.
Endemicity:
Endemic to Western Ghats.
84
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Clockwise from left: Fresh specimen UN by Raju Kasambe; Fresh specimen UN by Raju
Kasambe and old specimen UN by Ravi Vaidyanathan
Distribution:
P. d. demoleus Linnaeus, 1758: Throughout India below 2000 m elevation.
85
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution: Subspecies:
P. p. tamilana Moore, 1881: Western Ghats as far north as Maharashtra.
86
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution:
Peninsular India as far north as West Bengal.
87
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
UP by Vengolis UN by Vengolis
Distribution:
Western Ghats as far north as Goa.
Endemicity:
Endemic to Western Ghats.
88
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution: Subspecies:
P. a. aristolochiae (Fabricius, 1775): Throughout India.
89
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution:
Western Ghats south of Goa.
Endemicity:
Endemic to Western Ghats.
90
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution:
Peninsular India to W. Bengal. Straggler to the Andamans and Uttarakhand.
91
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Papilionidae 2016
Distribution:
Maharashtra to Kerala.
Endemicity:
Endemic to Western Ghats.
92
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Family Facts
Family Pieridae: Whites and Yellows
Most butterflies of the Family Pieridae are white or yellow in color, as the family's
common name suggests. They have black, red or orange markings.
Eggs are generally round and elongated, like a cylinder, and ribbed. They are laid
either singly or in batches. Caterpillars are generally green and without hairs or
spines. Pupae typically have both a cremaster and a silk girdle, and often are
pointed at the head. They are anchored at the tail.
93
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
P. c. canis Evans, 1912: Kerala; Tamil Nadu.
94
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
Throughout India except north eastern states.
95
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Clockwise from top left: Male UN, Male UP, Female UP and Female UN all by Raju Kasambe
Distribution: Subspecies:
C. n. evagete (Cramer, [1779]): Andhra Pradesh, Chhattisgarh, Delhi, Goa,
Gujarat, Haryana, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra,
Odisha, Rajasthan, Tamil Nadu, W. Bengal.
96
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UN by Aditya Joshi
Distribution: Subspecies:
C. n. remba (Moore, [1858]) Gujarat to Kerala.
97
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Upper- UP Male; Lower: UP Female; Right side: UN. All by Raju Kasambe
Distribution:
Throughout India excluding the North Eastern states and Jammu & Kashmir.
98
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Upper- UP Male; Lower: UN; Right side: UP Yellow form. All by Raju Kasambe
Distribution: Subspecies:
I. p. sesia (Fabricius, 1777): Throughout mainland India.
99
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
Throughout India except Andaman & Nicobar Is., Lakshadweep.
100
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
Maharashtra to Kerala.
101
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UN by Rahul K. Natu
Distribution:
A. l. lalage (Doubleday, 1842): Uttarakhand to N.E. India; Kerala, Tamil Nadu.
102
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
A. i. shiva (Swinhoe, 1885): Goa, Karnataka, Kerala, Tamil Nadu.
103
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Clockwise from top left: Female UN and Female UP, Male UP and Male UN all by Raju
Kasambe
Distribution: Subspecies:
A. a. darada (C. & R. Felder, [1865]): Uttarakhand to N.E. India; Maharashtra to
Kerala, Odisha.
Subspecies:
A. a. swinhoei (Moore, 1905): Gujarat to Kerala; Madhya Pradesh, Odisha.
105
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
A. l. latifasciata Moore, 1881: Maharashtra to Kerala.
106
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
L. n. nina (Fabricius, 1793): Throughout India east of Punjab, including the
Andaman Is.
107
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
Subspecies: H. g. australis Butler, 1898: Gujarat to Kerala.
108
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
C. a. amata (Fabricius, 1775): Delhi; Gujarat; Haryana; Maharashtra; Punjab;
Rajasthan; Uttar Pradesh.
Subspecies:
C. a. modestus (Butler, 1876): Andhra Pradesh; Karnataka; Kerala; Odisha; Tamil
Nadu; W. Bengal.
109
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
C. p. vestalis (Butler, 1876): Delhi, Haryana, Gujarat, Punjab, Rajasthan, Uttar
Pradesh.
110
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
C. f. fausta (Olivier, 1804): Delhi, Haryana, Gujarat, Madhya Pradesh,
Maharashtra, Punjab, Rajasthan, Uttar Pradesh.
Subspecies:
C. f. fulvia (Wallace, 1867): Karnataka, Kerala, Tamil Nadu.
111
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
C. e. etrida (Boisduval, 1876): Andhra Pradesh, Delhi, Goa, Gujarat, Haryana,
Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Punjab,
Rajasthan, Tamil Nadu, Uttar Pradesh, Uttarakhand.
112
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Clockwise from left: UN, UP Male and UP Female all by Dattaprasad Sawant
Distribution:
C. a. aurora (Cramer, [1780]): Goa; Gujarat; Karnataka; Kerala, Madhya Pradesh,
Maharashtra, Tamil Nadu.
113
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
C. d. danae (Fabricius, 1775): Goa, Gujarat, Karnataka, Kerala, Madhya Pradesh,
Maharashtra, Tamil Nadu.
114
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UP by Saish Borkar
Distribution:
Subspecies: P. c. ceylanica (C. & R. Felder, 1865): Goa to Kerala.
115
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Left: UP female Philomela form by Dr. Amol Patwardhan and right UN Male by Raju Kasambe
Distribution:
P. v. hippia (Fabricius, 1787): Throughout India except Jammu & Kashmir, Punjab
and Rajasthan.
116
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
Throughout India.
117
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UN by Raju Kasambe
Distribution:
Throughout India.
118
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UN by Raju Kasambe
Distribution:
E. b. rubella (Wallace, 1867): Throughout India including the Andaman and
Nicobar Is.
119
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
E. l. laeta (Boisduval, 1836): Throughout India, west of Sikkim and West Bengal.
120
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UN by Raju Kasambe
Distribution: Subspecies:
E. a. shimai Yata & Gaonkar, 1999: Karnataka; Kerala; Tamil Nadu.
121
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution:
E. h. hecabe (Linnaeus, 1758): Throughout India, including the Andaman and
Nicobar Islands.
122
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
Distribution: Subspecies:
E. b. davidsoni Moore, 1906: Gujarat to Kerala.
123
Butterflies of Western Ghats: Dr. Raju Kasambe Family:: Pieridae 2016
UN by Anila Manalil
Distribution:
Kerala; Tamil Nadu.
124
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Family Facts
Lycaenidae: Blues
Most butterflies belonging to the family Lycaenidae has blue uppersides. The
butterflies are generally characterized by: 1) small size, 2) reduced forelegs in
males but full-sized forelegs in females; thus they use only the remaining four legs.
Females have normal six legs. 3) A slightly different pattern of wing veins.
Many of the blues have lobes to their hind wings. Many have short or long fluffy
tails to their hind wings.
Eggs generally appear round and flattened like turbans. Caterpillars generally are
small, shaped like slugs, and hairy. The caterpillars of many of the species of
blues have a dorsal secretory organ which produces a sugary solution which
attracts to ants. The ants feed on the solution and in turn protect the caterpillar
from predators (a symbiotic behavior referred to as "tending"). Pupae are
generally small and round, may have a silk girdle, and are located near or on the
ground. Generally no cocoon is formed.
125
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
A. b. suffusa Moore, 1882. Gujarat southwards to Karnataka and Tamil Nadu and
eastwards to West Bengal; Himachal Pradesh to N.E. India.
Note: The similar looking Plum Judy Abisara echerius (Stoll, 1790) (Syn. Straight
Plum Judy) (Subspecies: A. e. prunosa Moore, 1879) is distributed across Kerala
and Tamil Nadu in Southern Western Ghats. It has a Wing span of 40–50 mm. It’s
larval host plants are Ardisia spp., Maesa indica and Embelia robusta, R. laeta
(family Primulaceae).
126
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
L. d. massalia Doherty, 1891. Karnataka. Sikkim to N.E. India.
127
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
S. e. epius (Westwood, 1852): Uttarakhand to N.E. India; Gujarat to Kerala and
east to W. Bengal.
128
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
T. n. nyseus (Guerin-Meneville, 1843): Maharashtra to Kerala, eastward to Andhra
Pradesh; Himachal Pradesh; Uttarakhand; Uttar Pradesh; Delhi.
129
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
C. r. rosimon (Fabricius, 1775): Throughout India including Andaman & Nicobar
Is.
130
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Maharashtra to Kerala; Sikkim to N.E. India.
131
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
C. d. decidia (Hewitson, 1876): Peninsular India; Sikkim to N.E. India.
132
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Subspecies: D. e. ethion (Westwood, 1851): Andaman & Nicobar Is. (Andamans);
Gujarat to Kerala; Uttarakhand to N.E. India.
133
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Goa to Kerala; Himachal Pradesh to West Bengal and Chhattisgarh.
134
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Throughout India excluding Jammu & Kashmir; Rajasthan and N.E. India.
135
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
T. b. nigra Bethune-Baker, [1918]: India north of Maharashtra to West Bengal.
136
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Throughout India except Jammu & Kashmir.
137
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Throughout India except the N.E. states.
138
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Throughout India except the N.E. states.
139
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
A. j. gamra (Lederer, 1855): India except east of W. Bengal.
140
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
141
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
M. m. thwaitesi Moore, [1881]: Maharashtra to Kerala; Sikkim; South India toWest
Bengal. Single record from Arunachal Pradesh.
142
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
A. p. felderi Toxopeus, 1927: Gujarat to Kerala.
143
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Dhaval Momaya
Distribution:
U. a. mavisa (Fruhstorfer, 1917): Karnataka, Kerala and Tamil Nadu.
144
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Anila Manalil
Distribution: Subspecies:
C. l. lavendularis (Moore, 1877): Karnataka; Kerala and Tamil Nadu.
145
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Clockwise from left: WSF UN and DSF UN by Raju Kasambe. UP Female by Sneha
Distribution:
C. l. lajus (Stoll, [1780]): Throughout India.
146
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
E. l. syntala Cantlie, 1963: Gujarat southwards to Andhra Pradesh and Kerala.
147
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
C. p. parrhasius (Fabricius 1793): Rajasthan to Kerala; eastwards to Uttar Pradesh;
Himachal; Pradesh and Uttarakhand.
148
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
149
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Clockwise from left: UN by Raju Kasambe, UP male by Jeevan Jose, UP female by Raju
Kasambe
Distribution:
Z. o. indica (Murray, 1874): Throughout India as far east as Jharkhand.
150
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
P. m. ossa (Swinhoe 1885): Maharashtra to Kerala and Andhra Pradesh.
151
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Throughout India; Andaman and Nicobar Is..
152
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
S. India; N. India from Punjab to N.E. India.
153
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Clockwise from left: UN by Raju Kasambe, Male UP by Jeevan Jose, Female Up by Anila
Manalil
Distribution:
Throughout India.
154
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Throughout India.
155
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Rohith Sanjay
Distribution: Subspecies:
A. e. emolus (Godart, 1824): Bihar; Sikkim; Maharashtra southwards to Kerala and
eastwards to West Bengal and N.E. India.
156
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
A. l. lycaenina (Felder, 1868): Gujarat southwards to Kerala and eastwards to
Odisha and W. Bengal.
157
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
C. s. strabo (Fabricius, 1793): Throughout India including Andaman & Nicobar Is.
158
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
C. p. exiguus (Distant, 1886): Karnataka to Kerala; Sikkim to N.E. India.
159
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Throughout India including Andaman & Nicobar Is.
160
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
J. b. bochus (Stoll, [1882]): Andaman & Nicobar Is. (Andamans); throughout
India.
161
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
J. c. aelianus (Fabricius, 1793): Gujarat south to Kerala and eastward to West
Bengal; Uttarakhand to N.E. India.
162
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Ashok Sengupta
Distribution: Subspecies:
N. p. continentalis Fruhstorfer, 1916: Sikkim to N.E. India; Maharashtra to Kerala.
163
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Subspecies: N. k. canaraica Toxopeus,1927: Gujarat to Kerala.
164
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
P. n. nora (C. Felder, 1860): Throughout India except arid regions.
165
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
P. d. indica (Evans, [1925]): India including Andaman & Nicobar Is. (Andamans).
166
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Divakar Thombre
Distribution:
P. n. hampsoni (de Niceville, 1885): Peninsular India as far north as Maharashtra;
Uttarakhand to N.E.India.
167
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Uttarakhand to N.E. India; Maharashtra to Kerala; Jharkhand and Andaman Is.
168
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
C. t. thetis (Drury, 1773): Gujarat east to Odisha and south to Kerala.
169
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Vengolis
Distribution:
Goa to Kerala.
170
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Clockwise from left: UN and UP Male by Raju Kasambe; UP Female by Ravi Vaidyanathan
Distribution:
C. a. dentata Moore, 1879: Himachal Pradesh to N.E. India; Gujarat to Kerala;
Madhya Pradesh to Odisha.
171
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Divakar Thombre
Distribution: Subspecies:
I. t. arsaces Fruhstorfer, 1907: Gujarat to Madhya Pradesh and Kerala.
172
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Clockwise from left: UN and Female UP by Raju Kasambe; Male UP by Shyam Ghate
Distribution: Subspecies:
A. n. dina Fruhstorfer, 1907: Gujarat to Kerala and W. Bengal; Assam.
173
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Prashanth Bhat
Distribution:
T. m. kanara Evans, 1925: Maharashtra to Kerala.
174
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
A. a. amantes (Hewitson, 1862): Gujarat to Andhra Pradesh and southwards to
Kerala.
175
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Uajith
Distribution:
A. a. indicus Riley, 1923: Karnataka; Kerala; Tamil Nadu; Uttarakhand to N.E.
India.
176
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by J.M. Garg
Distribution: Subspecies:
A. c. pirama (Moore, [1881]): Maharashtra to Kerala.
177
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Anila Manalil
Distribution:
A. b. bazaloides (Hewitson, 1878): Sikkim to N.E. India, Maharashtra to Kerala.
178
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
S. q. quercetorum (Moore, [1858]): Uttarakhand to N.E. India.
179
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Anila Manalil
Distribution:
Z. t. todara (Moore, [1884]): Goa to Kerala.
180
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
L. a. atymnus (Stoll, 1780): Maharashtra and Madhya Pradesh to Kerala;
Uttarakhand to West Bengal &. N. E. India.
181
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
S. v. vulcanus (Fabricius, 1775): Throughout India.
182
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
S. l. lazularia (Moore, [1881]): Maharashtra to Kerala.
183
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Gujarat to Kerala.
184
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Maharashtra to Tamil Nadu.
UN by Raju Kasambe
185
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
S. i. ictis (Hewitson, 1865): Rajasthan northwards to Himachal Pradesh, eastwards
to W. Bengal and southwards to Kerala.
186
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
S. e. elima (Moore, 1877): Gujarat eastwards to W. Bengal and southwards to
Kerala; Himachal Pradesh to N.E. India.
187
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Makarand Kulkarni
Distribution:
Maharashtra to Kerala; Uttarakhand; Uttar Pradesh to N.E. India.
188
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
P. d. deva (Moore, 1858): Madhya Pradesh; Kerala to Maharashtra; West Bengal.
189
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Pinakin Karve
Distribution: Subspecies:
A. b. sudica (Evans, 1926): Maharashtra to Kerala.
190
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
T. c. cippus (Fabricius, 1798): Andaman & Nicobar Is. (Andamans); throughout
India except arid regions.
191
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Sikkim to N.E. India, Tamil Nadu, Karnataka, Kerala.
192
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Goa to Kerala; Uttarakhand to N.E. India.
193
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Praveen G Nair
Distribution:
Subspecies: E. j. macanita Fruhstorfer, 1912: Goa to Kerala.
194
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
C. f. butleri Cowan, 1965: Maharashtra to Kerala.
195
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Kerala to N.E. India.
196
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
H. o. cingalensis Moore, 1884: Maharashtra to Kerala.
197
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
S. India, Himachal Pradesh to N.E. India.
198
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Hemant Ogale
Distribution: Subspecies:
C. m. callone Fruhstorfer, 1915: Maharashtra to Kerala.
199
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Maharashtra to Kerala; Uttarakhand to N.E. India; Andaman Is.
200
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by R. Ravi
Distribution:
Karnataka; Kerala; Tamil Nadu.
201
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Goa to Kerala; Sikkim to N.E. India.
202
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Clockwise from left: UN by Raju Kasambe, Female UP by Anila Manalil and Male UP by
Divakar Thombre
Distribution: Subspecies:
D. e. epijarbas (Moore, 1857): South India to West Bengal.
203
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Throughout India.
204
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
V. p. ghela Fruhstorfer, 1912: Maharashtra to Kerala.
205
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Aditya Joshi
Distribution: Subspecies:
R. v. lazulina (Moore, 1879): Maharashtra to Kerala, West Bengal.
206
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
R. m. schistacea (Moore, 1879): Throughout India; Andaman & Nicobar Is.
(Andamans).
207
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution: Subspecies:
R. i. sorya (Kollar, [1844]): Jammu & Kashmir; Odisha; Punjab; Peninsular India;
Uttarakhand; Uttar Pradesh and W. Bengal.
208
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
Distribution:
Karnataka, Kerala, Tamil Nadu.
209
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Lycaenidae 2016
UN by Raju Kasambe
Distribution:
Subspecies: B. p. moorei Fruhstorfer, 1904: Goa to Kerala.
210
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Family Facts
Family Nymphalidae: Brush-footed Butterflies
The family is known as the Brush-footed butterflies because the forelegs of the
adults are small and hairy resembling tiny brushes, and are not used for
walking.
The butterflies in this family vary considerably in their appearance in all stages
of the life cycle (adult, eggs, larval, pupal), but generally can be characterized
by the following, 1) size of the forelegs is reduced (except female of Beaks),
thus they use only four legs to walk or perch; 2) medium to large in size and
brightly and/or uniquely marked; 3) the pattern of wing veins of the forewing is
unique; and 4) the rigid antennae have club shaped tips.
Eggs vary in shape and in their arrangement on the plant. Caterpillars vary
considerably in their appearance, but are often hairy or spiny. Pupae have a
cremaster from which they are suspended upside down, but have no silk girdle
and form no cocoon.
211
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Dhaval Momaya
Distribution: Subspecies:
L. l. lepitoides Moore, 1901: Gujarat to Kerala.
212
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
L. m. rama Moore, 1872: Maharashtra to Kerala; Andhra Pradesh.
213
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
P. a. aglea (Stoll, [1782]): Gujarat to Kerala; Chhattisgarh; Odisha; W. Bengal.
214
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Kishen Das
Distribution:
Karnataka; Kerala; Tamil Nadu.
215
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
T. s. dravidarum (Fruhstorfer, 1899): Gujarat east to Odisha and south to
Kerala.
216
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
T. l. exoticus (Gmelin, 1790): Throughout India including Lakshadweep;
Andaman & Nicobar Is.
217
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
D. c. chrysippus (Linnaeus, 1758): Throughout India.
218
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
D. g. genutia (Cramer, [1779]): Throughout India.
219
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
E. c. core (Cramer, [1780]): Throughout India.
220
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
E. s. coreta (Godart, 1819): Peninsular India.
221
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
E. k. kollari C. & R. Felder, [1865]: Gujarat eastwards to W. Bengal and
Odisha.
222
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
I. m. kanarensis (Moore, 1890): Maharashtra to north Karnataka.
Subspecies:
I. m. malabarica (Moore, 1877): Southern Karnataka to Kerala.
223
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
Ch. p. imna Butler, 1870: Peninsular India as far north as Gujarat; Madhya
Pradesh; Odisha to N. E. India.
Note: The similar looking Tawny Rajah Charaxes bernardus (Fabricius, 1793)
is found in Sikkim to N.E. India; Uttarakhand.and Andaman Is.
224
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
Ch. s. solon (Fabricius, 1793): Rajasthan to Kerala; Delhi; Himachal Pradesh to
Sikkim and W. Bengal.
225
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
P. a. athamas (Drury, [1773]): Himachal Pradesh to N.E. India; peninsular India
south of Gujarat and Jharkhand.
226
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Aditya Joshi
Distribution: Subspecies:
P. s. wardii (Moore, 1896): Maharashtra to Kerala.
227
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raju Kasambe
Distribution:
P. a. agraria (Swinhoe, 1887): Gujarat to Madhya Pradesh and Kerala;
Himachal Pradesh to N.E. India.
228
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raghu
Distribution:
D. l. lepida Moore, 1857: Goa to Kerala.
229
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Richard Parker
Distribution:
A. p. friderici Fruhstorfer, 1904: Kerala.
230
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
M. a. anaxias Hewitson, 1862: Karnataka to Kerala.
231
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Clockwise from left: WSF UN by Raju Kasambe, UP by Ms. Marvelyn Dias, DSF UN by
Shyam Ghate
Distribution: Subspecies:
M. v. subdita (Moore, [1890]): Kerala to Odisha.
Subspecies:
M. v. visala Moore, [1858]: Uttarakhand to N.E. India; Gujarat to ?Goa and
Madhya Pradesh to W. Bengal.
232
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Anila Manalil
Distribution:
Subspecies: H. m. davisonii (Moore, [1891]): Kerala and Tamil Nadu.
233
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Kerala and Tamil Nadu.
234
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Endemicity:
Endemic to Western Ghats and Sri Lanka.
Distribution:
M. p. junonia Butler, 1868: Goa to Kerala.
235
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
M. p. tabitha (Fabricius, 1793): Peninsular India south of the Himalaya to
Kerala.
236
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
M. m. polydecta (Cramer, [1777]): Peninsular India from Gujarat to W. Bengal
to Kerala; Lakshdweep.
237
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
L. r. neelgheriensis (Guerin-Meneville, 1843): Rajasthan east to West Bengal
and south to Kerala.
238
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Vinayaraj
Distribution:
L. d. todara Moore, 1881: Goa to Kerala; Chhattisgarh and Odisha..
239
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
L. e. ragalva Fruhstorfer, 1911: Gujarat eastwards to Odisha and southwards to
Kerala.
240
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raju Kasambe
Distribution:
Y. a. mahratta Moore, 1884: Throughout India.
241
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
Y. b. madrasa Evans, 1924: Gujarat to Kerala.
242
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Y. c. ceylonica Hewitson, 1865: Goa eastwards to Odisha and southwards to
Kerala.
243
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Aditya Joshi
Endemicity:
Endemic to Western Ghats (Karnataka southwards).
Distribution:
Karnataka and Tamil Nadu.
244
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Y. h. huebneri Kirby, 1871: Throughout India.
245
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Karnataka to Kerala.
246
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
O. m. mandata (Moore, 1857): Maharashtra; Madhya Pradesh and Chhattisgarh
south to Kerala.
247
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
M. l. leda (Linnaeus, 1758): Throughout India including Lakshadweep and
Andaman & Nicobar Is.
248
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
M. ph. varaha Moore, 1857: Maharashtra to Kerala.
249
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Raju Kasambe
Distribution: Subspecies:
M. z. gokala Moore, 1857: Maharashtra to Kerala.
250
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by Subhiksha S.
Endemicity:
Endemic to Western Ghats.
Distribution:
Karnataka to Kerala.
251
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
E. h. caudata Butler, 1871: Maharashtra to Kerala.
252
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
V. e. saloma de Niceville, 1886: Maharashtra to
Kerala.
253
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Endemicity:
Endemic to Western Ghats and Sri Lanka.
Distribution:
C. t. thais (Fabricius, 1787): Gujarat to Kerala.
254
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
C. e. maja Fruhstorfer, 1898: Maharashtra to Kerala.
255
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
P. a. mercea (Evans, 1924): Maharashtra to Kerala.
256
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
P. p. phalantha (Drury, [1773]): Throughout India.
257
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
A. h. castetsi Oberthur, 1891: Kerala; Tamil Nadu.
Subspecies:
A. h. hybrida Evans, 1912: Tamil Nadu.
Subspecies:
A. h. hyperbius (Linnaeus, 1763): Rajasthan to Madhya Pradesh, Gujarat, Uttar
Pradesh; Jammu & Kashmir to N.E. India.
258
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Endemicity:
Endemic to Western Ghats and Sri Lanka.
Distribution:
C. n. mahratta Moore, 1872: Maharashtra to Kerala.
259
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Throughout India; Lakshadweep Is.
260
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
M. p. undifragus Fruhstorfer, 1906: Gujarat eastwards to Odisha and
southwards to Kerala.
261
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
A. p. perius (Linnaeus, 1758): Kerala north to Maharashtra, Madhya Pradesh
and Jharkhand; Himachal Pradesh to N.E. India.
262
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
A. r. karwara Fruhstorfer, 1906: Karnataka to Kerala.
263
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
A. s. kanara (Evans, 1924): Goa east to Jharkhand and southwards to Kerala.
264
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
A. n. inara (Westwood, 1850): Uttarakhand to N.E. India; Karnataka to Kerala
and northwards to Odisha.
265
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
P. h. hordonia (Stoll, [1784]): Maharashtra eastwards to W. Bengal and
southwards to Kerala; Uttarakhand to N.E. India.
266
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Rajkamal Goswami
Distribution: Subspecies:
P. s. davidsoni Eliot, 1969: Uttarakhand to N.E. India, peninsular India south of
Maharashtra.
267
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
N. h. varmona Moore, 1872: Gujarat, Madhya Pradesh and Jharkhand
southwards to Kerala.
268
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
Ph. c. nilgirica (Moore, 1889): Gujarat eastwards to W. Bengal and southwards
to Kerala.
269
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
N. j. jumbah Moore, [1858]: Gujarat eastwards to W. Bengal and southwards to
Kerala; Sikkim to .N.E. India.
270
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
L. v. kanara (Evans, 1924): Goa to Kerala.
271
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Raju Kasambe
Distribution: Subspecies:
N. n. hampsoni Moore, 1899: Goa to Kerala.
272
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Raju Kasambe
Distribution: Subspecies:
N. c. kallaura Moore, 1881: Maharashtra to Kerala.
273
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Jeevan Jose
Distribution: Subspecies:
P. s. virens Moore, [1877]: Maharashtra to Kerala.
274
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
E. a. anagama Fruhstorfer, 1913: Maharashtra to Odisha; Himachal Pradesh to
Uttarakhand and Uttar Pradesh.
Subspecies:
E. a. meridionalis Fruhstorfer, 1913: Maharashtra to Kerala and Andhra
Pradesh.
275
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
E. l. arasada Fruhstorfer, 1913:
Maharashtra to Kerala.
Subspecies:
E. l. lubentina (Cramer, [1777]): Maharashtra and Gujarat eastwards to
Haryana, Odisha andWest Bengal; Himachal Pradesh to N.E. India.
276
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Tamil Nadu to Gujarat and Rajasthan, eastwards to W. Bengal and along the
Himalaya from Uttarakhand to W. Bengal.
277
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
C. t. telchinia (Menetries, 1857): Karnataka; Sikkim to N.E. India.
278
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Raju Kasambe
Distribution: Subspecies:
C. l. miyana (Fruhstorfer, 1913): Maharashtra eastwards to Odisha and
southwards to Kerala.
279
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Clockwise from left: Female UN, Male UP and Female UP all photos by Hemant Ogale
Distribution: Subspecies:
E. e. laudabilis Swinhoe, 1890: Gujarat to Kerala.
280
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Nandish Songire
Distribution: Subspecies:
C. t. indica Evans, 1924: Gujarat to Kerala.
281
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP by Raju Kasambe
Distribution: Subspecies:
A. a. indica (Moore, 1884): Peninsular India from Gujarat and W. Bengal to
Kerala.
282
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
A. m. merione (Cramer, [1777]): Gujarat to Kerala and Andhra Pradesh.
283
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Gujarat eastwards to Chhattisgarh and southwards to Tamil Nadu.
284
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
R. p. atacinus Fruhstorfer, 1913: Kerala to Maharashtra.
285
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UN by PKG Mohan
Distribution: Subspecies:
E. c. meridionalis Wood -Mason, 1881: Maharashtra to Kerala.
286
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
V. i. pholoe Fruhstorfer, 1912: Karnataka to Kerala.
287
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Throughout India; straggler on Andamans and Lakshadweep Is.
288
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
UP and UN by L. Shyamal
Distribution: Subspecies:
K. c. viridis Evans, 1924: Karnataka to Kerala.
289
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
J. a. atlites (Linnaeus, 1763): Throughout India except arid regions; Andamans
and Nicobars.
290
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
J. a. almana (Linnaeus, 1758): Throughout India including Andaman Is.
291
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
J. h. hierta (Fabricius, 1798): Throughout India except N.E. India and the
Andaman Is.
292
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
J. i. pluviatalis Fruhstorfer, 1900: Peninsular India to Madhya Pradesh.
293
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
J. l. vaisya (Fruhstorfer, 1912): Rajasthan to Kerala and eastwards to Jharkhand.
294
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Clockwise from left: Male UP by J.M. Garg, Female UP and UN by Raju Kasambe
Distribution: Subspecies:
J. o. swinhoei Butler, 1885: Jammu & Kashmir to Kerala and W. Bengal.
295
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Clockwise from top left: Female UP, Female UN, Male UP and Male UN by Raju Kasambe
Distribution:
H. b. jacintha (Drury, 1773): Throughout India except very arid regions;
Andaman & Nicobar Is.
296
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
297
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
K. i. huegeli (Kollar, [1844]): Jammu & Kashmir to Uttarakhand; Jharkhand,
Eastern Ghats, Madhya Pradesh and Gujarat.
298
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution:
Maharashtra to Kerala.
299
Butterflies of Western Ghats: Dr. Raju Kasambe Family: Nymphalidae 2016
Distribution: Subspecies:
D. b. malabarica Fruhstorfer, 1899: Goa to Kerala.
300
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Butterfly garden is a garden where you can see lot of butterflies belonging to
different species at one place and in good numbers. An ideal butterfly garden is
nothing but a miniature representation of the forest in the adjoining area where
various plants and flowers are grown. The environment is made as conducive as
possible for butterflies found in the area.
To set up a new butterfly garden what we need is a suitable piece of land at the
right location, man power, working knowledge of landscaping, good knowledge
about butterflies and their requirements, knowledge about plants and finances to
run the garden. If you want to make it a public place, you need some
advertisement or selling skills to attract people to visit the butterfly garden.
Butterfly garden can be a closed one or an open one. A closed butterfly garden
can is an entire garden covered with green nets. The butterflies are released into
this enclosed area either as adult butterflies or are reared in the garden itself. In
Singapore and London butterfly parks, they procure pupa (chrysalis) and keep
them in small boxes till the butterflies emerge. As soon as the butterflies emerge
they are released into the garden. In absence of any predators, the butterflies
survive till they die naturally.
301
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
But the Indian Wildlife Protection Act (1972) considers butterflies as wild
animals and it is illegal to keep any wild animal in an enclosure (or captivity, in
the legal language). As soon as there is an enclosure, it becomes a zoo. And to
run a zoo, permission is required from Central Zoo Authority of India and also
it is mandatory to follow their guidelines for establishing and managing a zoo.
Getting permission from this authority (though creating a butterfly park may be
a good objective) is ‘mission impossible’ for a general citizen.
302
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
It is important to have a good location for the butterfly garden, especially the
open butterfly garden. The best location will be near a patch of forest, as the
butterflies can be attracted to the garden by fulfilling the necessary
requirements. However, it is not necessary to have big plot to attract butterflies
to the garden. Even a small plot can be converted into a decent butterfly garden.
It is better to avoid setting up a new butterfly garden in very dry areas or very
cold areas, deserts, and in highly urbanized areas with not many butterfly
species around.Once the land is acquired, it is important to do landscaping of
the plot as per the requirements of a butterfly garden. If it already has lot of tree,
there is no need to cut the tree, but to ensure plantation of new plants at the right
locations. Landscaping should be in such way that there are places which
provide shade, lot of sunshine, and wet patches too.
303
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Once the location of the butterfly garden is decided, we need to know what are
the requirements of butterflies or what are the things which attract them? As in
India, we cannot have a closed butterfly garden, the best option we have is to
attract the butterflies and make them stay around in the butterfly garden.
Butterflies will surely stay around in the garden, if we fulfil all their
requirements in the garden itself.
As mentioned above the life cycle of a butterfly completes in four stages, and
each butterfly species lays its eggs on a specific plant (or a choice of few
species of plants). The larva (or caterpillars) feed on these plants and hence
these plants are termed as larval host plants (LHP). For example the Common
Rose butterfly lays its eggs on Aristolochia indica, Spot Swordtail, Common
Jay and Tailed Jay lays their eggs on Polyalthia longifolia, Commom Mormon
lays its eggs on Murraya koenigii (Curry Leaf) and Citrus aurantifolia (Lime
tree).
The caterpillar which feeds on the LHP metamorphoses into a pupa in due
course of time. The pupa is generally well camouflaged and stays immobile till
an adult butterfly emerges out of it (there is nothing like a ‘baby butterfly’).
The more is the diversity of the larval host plants in the butterfly garden the
more number of butterfly species will start breeding in the garden. And there are
more chances of the butterflies staying back in the area if they can fulfil all their
requirements in the area. Hence, as a part of developing the butterfly garden, it
is a continuous process to find out more and more larval host plants and plant or
grow them in the garden. It is important to have better understanding among the
staff (at least the gardener) of the butterfly garden to prevent uprooting of the
larval host plants, mistaking them to be useless weeds. This is important in view
of the fact that some butterfly species lay their eggs on grasses.
Butterflies do not have teeth they cannot eat solid food. Butterflies can only sip
liquid food with the help of a very thin tongue, called proboscis. And they do
not grow once they are borne, they do not need a diversity of food for physical
growth. What they need is liquid food which is rich in energy and acts like fuel
304
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
love the nectar of flowers. These butterflies are attracted to the flowers due
to their bright colours. Hence it is necessary to plant plots of flowering plants in
the garden. The flowering plants should be selected carefully in such a way that
throughout the year the garden has some plants flowering. Some of the common
plants which attract lot of butterfly species for nectaring are Lantana spp.,
Jamaican Blue Stachytarphaeta spp., Cockscomb Celosia spp., wild Xenia spp.
and Ixora species. A small herb Tridax indica attracts lot of blue (Lycaenid)
butterflies for nectaring.
Every butterfly has its own choice of flowers it visits, due to the fact that the
length of their proboscis varies
in different species. Butterflies
with short proboscis may not be
able to sip nectar from flowers
with a long tubular corolla.
Some skippers have extremely
long proboscis and hence can
sip nectar from flowers with
long tubular corolla. Many
large sized swallowtail
butterflies are not able to sip
nectar from very small flowers Silver Coxcomb Celosia argentea bushes attract many butterflies like
Pansies by Raju Kasambe
305
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
as the flowers cannot bear their weight. Hence some of the swallowtails do not
land on the flowers and keep fluttering while nectaring. Thus we need to a have
a diversity of flowering plants in the garden to cater to the needs of the various
species of butterflies. Also while planting the flowering plants, they should be
planted keeping in mind their expected height to which they will grow. This will
provide a vertical dimension to the garden. Small plots of flowering plants of a
particular species should be planted, thus adding to the aesthetic value of the
butterfly garden, besides attracting hordes of butterflies.
It is a wrong notion that all butterflies love nectar in flowers. There are many
species of butterflies (many belong to the brush-footed and brown butterflies)
which never visit a flower. These butterflies like to get their stock of food from
rotten fruits, decaying fish, crabs, or prawns, the scat or dung or urine of wild
animals and so on. These butterflies locate the food because of the strong smell.
Some of the butterflies which are attracted to these include the Common Nawab
Polyura athamas, Anamolous Nawab Charaxes agrarius, Black Rajah
Charaxes solon, Tawny Rajah Charaxes bernardus, Blue Oakleaf Kallima
horsfieldii, Orange Oakleaf Kallima inachus, Common Baron Euthalia
aconthea, Gaudy Baron Euthalia lubentina, Common Evening Brown Melanitis
leda, Angled Castor Ariadne ariadne, Common Palmfly Elymnias hypermnestra
and many bushbrowns Mycalesis spp.
Overripe and rotten fruits are kept in butterfly garden to fulfil the requirement of butterflies which don’t like
flowers. This is from Butterfly Garden at Singapore Airport by Raju Kasambe
306
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Rotten fruits can be kept in feeding trays in the butterfly garden to attract these
butterflies. These feeding trays with rotting fruits may attract ants, which in turn
will disturb the butterflies. To manage this problem, each feeding tray should be
kept in another slightly larger tray filled with water. This will prevent ants to
reach the bowl with fruits, creating a moat-like situation like a. The feeding tray
can also be kept hanging in trees at various locations in the garden. Rotten or
overripe fruits of Pineapple Ananas comosus, Custard apple Annona reticulata,
banana, guava and Sapota (Sapodilla or chikoo) Manilkara zapota are useful in
attracting butterflies.
Males of some butterfly species need specific alkaloids for reproduction. These
alkaloids are provided by plants like Rattlepod Crotalaria, Turnsole
Heliotropium, and Eupatorium species. The male butterflies flock these plants
in huge numbers to suck these alkaloids and is a pleasant sight to see the
butterflies engrossed in the activity. They need these chemicals to synthesize
sex pheromones to attract females. Developing small plots of these plants
ensure flocks of butterflies especially the tiger butterflies namely, Blue Tiger
Tirumala limniace, Glassy Tiger Parantica aglea, Dark Blue Tiger Tirumala
Milkweed butterflies like Tigers and Crows are attracted towards Crotalaria to get alkaloids by Raju Kasambe
307
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Also, before planting the plots of plants, a permanent arrangement for watering
the plants should be done. This could be the sprinklers or other methods, which
will save lot of manpower in future needed to water the entire garden nearly
every day.
Many species of butterflies gather at wet muddy patches for mud-puddling by Raju Kasambe
308
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Insecticides and weedicides should not be sprayed in butterfly garden in any case
Another thing is avoiding plantation of rose plants. These are useless for
butterflies and need spaying of insecticides to protect it from various moth
caterpillars.
Avoid using chemical fertilizers. Instead, the garden can have a small place
where natural fertilizers can be produced using the leaf litter, cattle dung and
other garbage from the garden. This natural fertilizer should be used for the
larval host plants for their healthy growth.
Landscaping
While setting up an open butterfly garden, it is necessary to have a plan of
landscaping. But if there are trees already in place, the plan should be designed
accordingly without cutting the existing trees. The plan should clearly describe
the plots for flowering trees and take care of the needs of various species of
trees as per their requirement and dependence on sunlight. There should be a
small nursery totally enclosed to protect small saplings of various plants
collected for the garden. Few saplings of each larval host plant (at least the rare
once) should be protected here as genetic pool. Sometimes entire plants are
finished by hordes of caterpillars. This stock will help replenish the plants in the
garden again.
309
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Butterfly garden landscaping should take into consideration many factors like parking space, toilets, etc.
(Photo- Dr. Raju Kasambe)
Sunshine is very important in the life of butterflies as they are cold blooded
animals and need to bask in sunlight before they start their activities in the
morning. They because active after basking in sunlight for some time. Hence
the garden should have lot of flower beds with lot of sunshine.
While planting large trees in the garden, lot of planning and futuristic thinking
is needed. When they grow, they should not create shadow in entire butterfly
garden. If he garden is very big, the trees can be planted along the eastern edge
or they should be planted along the western edge or forming a line running
north-south in the middle of the garden. This will keep big areas with lot of
sunlight.
310
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
A small shade for visitors with resting facilities, a tea and snacks kiosk and
basic amenities like toilets ensure the comforts of the visitors.
It is important to maintain a genetic stock of all the larval host plants as well as
nectar plants in the form of seeds or grafts. The seeds should be collected
whenever they are found and kept safe till the pre-monsoon period. The seeds
should be grown in a netted nursery and then shifted to the butterfly garden.
This should be done at least to plant species Tagala and Aristolochia as the
caterpillars of Roses are known to finish the entire plants leaving nothing
behind.
Manpower
To manage an open butterfly garden you need to hire at least one gardener, a
plant expert, a butterfly expert and a person to manage the entire set up. Off
course, you can play few roles out of these.
You need some seeding money to start with butterfly gardening and there is no
guarantee that it will earn you money. If you have the land needed to set up the
butterfly garden available with you, the initial cost will be less. But there is
recurring cost to maintain a butterfly garden which includes purchase of
saplings, gardener, honorarium to butterfly and plant experts (many roles can be
played by one person), electricity bills, and water bills and so on. Butterfly
garden needs proper management and regular maintenance, otherwise the weeds
overgrow the larval host plants and soon it loses the beauty of being a ‘butterfly
garden’!
Anybody can rear a butterfly in his house! That’s pretty simple. You can search
for caterpillars on larval host plants of butterflies like a Lime tree Citrus spp. or
a Curry Leaf tree Murraya koenigii may have caterpillars of Common Mormon
Papilio polytes and Lime Butterflies Papilio demoleus. Pick that caterpillar and
bring with it a stock of tender leaves to feed it on. Keep it in a transparent box.
Keep some sticks in the box for the caterpillar to pupate. Clean the box
311
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Readymade butterfly pupae are brought and kept in specially designed ‘emergence cages’ in closed butterfly
gardens. The butterflies are released when they emerge from these pupae by Raju Kasambe
everyday for the excreta. The caterpillar will get converted to a pupa one day.
One fine morning you will notice that a beautiful butterfly has emerged from
the pupa. Free it as soon as it is ready to fly….It’s your baby after all!
In India, we can have small closed butterfly gardens in major zoos and at major
International Airports for recreation. For maintaining these gardens, we can start
rearing butterflies of belonging to the commoner but beautiful and colourful
species, not necessarily rare or those protected under various Schedules of the
Indian Wildlife Protection Act (1972), with the help of tribal communities and
create thousands of jobs.
Butterfly ‘nurseries’ for selected species (not listed under the various schedules
of the Act) can be set up where butterflies can be bred. The pupae reared from
such ‘nurseries’ can be supplied to butterfly parks around the country. This can
very well be done in villages around Western Ghats and the Himalayas by
setting up of big nurseries of local larval host plants in village lands or in
private lands. What is needed is little flexibility in the Indian Wildlife
Protection Act (1972) and initiative from the NGOs or forest department.
312
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Dos’:
1. Know the diversity of butterflies in your area.
2. Plant larval host plants.
3. Plant nectar plants.
4. Provide rotten fruits.
5. Keep a damp patch for mud puddling.
6. Keep some shady places.
Don’ts:
1. Avoid use of insecticides, weedicides (herbicides) or such chemicals.
Sammilan Shetty’s Butterfly Park, Santhadi House, Butterfly Park Road, Belvai
Village, Mangalore, Karnataka
313
Butterflies of Western Ghats: Dr. Raju Kasambe Butterfly Gardening 2016
Reference:
Kasambe, R. (2015) Butterfly Gardening in India: An Introduction. Research
Gate DOI: 10.13140/RG.2.1.4934.6164.=.
314
Butterflies of Western Ghats Endemic butterflies 2016
Papilionidae
Pieridae:
6. Nilgiri Clouded Yellow Colias nilagiriensis C. & R. Felder, 1859. Range:
Kerala and Tamil Nadu.
7. Nilgiri Grass Yellow Eurema nilgiriensis Yata, 1990. Range: Karnataka;
Kerala; Tamil Nadu.
8. Lesser Albatross Appias wardi (Moore, 1884). Range: Maharashtra to
Kerala.
Nymphalidae:
9. Travancore Evening Brown Parantirrhoea marshalli Wood-Mason, 1880.
Range: Karnataka to Kerala.
10. *Small Long-brand Bushbrown Mycalesis igilia Fruhstorfer, 1911. Note:
There are two sub-species, viz., M. i. igilia Fruhstorfer, 1911 ranging from
Karnataka to Kerala and M. i. mercea Evans, 1920 ranging from Madhya
Pradesh to Maharashtra)(Varshney & Smetacek (2015).
11. Pale-brand Bushbrown Mycalesis orcha Evans, 1920. Range: Karnataka to
Kerala.
315
Butterflies of Western Ghats Endemic butterflies 2016
12. Red-disc Bushbrown Mycalesis oculus Marshall, 1881. Range: Kerala and
Tamil Nadu.
13. Redeye Bushbrown Heteropsis adolphei (Guerin-Meneville, 1843). Range:
Karnataka; Tamil Nadu.
14. *Lepcha Bushbrown Mycalesis lepcha (Moore, 1880). Note: Not found in
Western Ghats (Varshney & Smetacek (2015).
15. Tamil Catseye Zipaetis saitis Hewitson, 1863. Range: Karnataka to Kerala.
16. Palni Fourring Ypthima yphthimoides (Moore, 1881). Range: Kerala and
Tamil Nadu.
17. Nilgiri Fourring Ypthima chenu (Guérin-Méneville, 1843). Range:
Karnataka and Tamil Nadu.
18. Southern Blue Oakleaf Kallima horsfieldi (Kollar, [1844]). Range:
Maharashtra to Kerala.
19. Nilgiri Tiger Parantica nilgiriensis (Moore, 1877). Range: Karnataka;
Kerala; Tamil Nadu.
20. Malabar Tree Nymph Idea malabarica (Moore, 1890). Range: Maharashtra
to Kerala.
Lycaenidae:
21. *Indian Pointed Pierrot Tarucus indicus Evans, 1932. Range: Dry zones
from Rajasthan south to Tamil Nadu and east to West Bengal; Assam. Note:
Listed as endemic but it is also reported from other areas.
22. Whitedisc Hedge Blue Celatoxia albidisca (Moore, 1884). Range:
Karnataka; Kerala and Tamil Nadu.
23. Kanara Oakblue Arhopala alea (Hewitson, 1862)(Syn. Rosy Oakblue).
Range: Goa to Kerala.
24. Abnormal Silverline Spindasis abnormis (Moore, [1884]): Range:
Maharashtra to Tamil Nadu.
25. Shiva’s Sunbeam Curetis siva Evans. 1954. Range: Goa to Kerala.
Hesperiidae:
26. *Spotted Small Flat Sarangesa purendra Moore, 1882. Treated as a species
in Gaonkar (1996). Listed as a sub-species Sarangesa purendra hopkinsi
316
Butterflies of Western Ghats Endemic butterflies 2016
Reference:
Roy, P K., Harikumaar, B., and Hassan, S. (2010): Kerala, an Authentic
Handbook. Information-Public Relations Dept., Thiruvananthapuram, Govt. of
Kerala.
317
Butterflies of Western Ghats Index of Common Names 2016
318
Butterflies of Western Ghats Index of Common Names 2016
Crow, Brown King 222 Emigrant, Mottled 118 Fourring, Nilgiri 244
Crow, Common Indian 220 Evening Brown, Dark 249 Fritillary, Indian 258
Cupid, Indian 147 Flash, Common Red 208 Hedge Blue, Plain 145
Cupid, Plains 155 Flash, Indigo 206 Hedge Blue, White 144
319
Butterflies of Western Ghats Index of Common Names 2016
Judy, Plum 126 Nymph, Malabar Tree 223 Pierrot, Balkan 136
Judy, Two-spot Plum 126 Nymphalidae 211 Pierrot, Banded Blue 133
320
Butterflies of Western Ghats Index of Common Names 2016
Royal, Peacock 191 Skipper, African Marbled 64 Tip, Great Orange 108
Royal, Silver 190 Skipper, Indian 68 Tip, Little (Small) Orange 112
Royal, Spotted 192 Sunbeam, Angled 171 Tip, Plain Orange 113
321
Butterflies of Western Ghats Index of Scientific Names 2016
322
Butterflies of Western Ghats Index of Scientific Names 2016
323
Butterflies of Western Ghats Index of Scientific Names 2016
324
Butterflies of Western Ghats Index of Scientific Names 2016
325
Butterflies of Western Ghats Important References 2016
Important References
Evans, W.H. (1932) The Identification of Indian Butterflies. (2nd Ed), Bombay Natural History Society,
Mumbai, India
Gaonkar, H. (1996) Butterflies of the Western Ghats, India, including Sri Lanka - A Biodiversity
Assessment of a Threatened Mountain System. Centre for Ecological Sciences, Bangalore, Zoological
Museum, Copenhagen, the Natural History Museum, London. Pp.82.
Kalesh, S & S K Prakash (2007) Additions of the larval host plants of butterflies of the Western Ghats,
Kerala, Southern India (Rhopalocera, Lepidoptera): Part 1. J. Bombay Nat. Hist. Soc. 104 (2): 235–238.
Kehimkar, I. (2008) The Book of Indian Butterflies. Bombay Natural History Society, Mumbai. Pp.497.
Kehimkar, I. (2016) BNHS Filed Guides - Butterflies of India. Bombay Natural History Society, Mumbai.
Pp.509.
Kunte, K. (2006) Additions to the known larval host plants of Indian butterflies. J. Bombay Nat. Hist. Soc.
103(1): 119–121.
Kunte, K. (2005) Butterflies of Peninsular India. Indian Academy of Sciences. Universities Press.
Pp.254+colour plates-31.
Roy, P K., Harikumaar, B., and Hassan, S. (2010) Kerala, an Authentic Handbook. Information-Public
Relations Dept., Thiruvananthapuram, Govt. of Kerala.
Wynter-Blyth, M.A. (1957) Butterflies of the Indian Region. Bombay Natural History Society, Mumbai,
India.
Varshney, R.K. & Smetacek, P. (eds.) (2015) A Synoptic Catalogue of the Butterflies of India. Butterfly
Research Centre, Bhimtal and Indinov Publishing, New Delhi, ii + 261 pp., 8 pl.
Important Websites:
http://www.ifoundbutterflies.org/
http://flutters.org/
326
Butterflies of Western Ghats: Dr. Raju Kasambe About the Book 2016
Abbreviations used:
DSF: Dry Season Form
WFS: Wet Season Form
UP: Upperside
UN: Underside
327
About author
Academic Qualification:
B.Sc., B.A., M.A. (Eng), D.B.M., M.Sc., M.Phil. (Env.),
B.Ed., Ph.D.
Job:
Working at Bombay Natural History Society (BNHS),
Mumbai since 2010 as Important Bird Areas (IBA)
Programme Manager and In-charge of Conservation
Education Centre (CEC), Goregaon East, Mumbai
Research publications
More than 100 research papers in international and national journals about birds and
butterflies.
Books Published
Indian Grey Hornbill (Lambert, Germany) (2011)
Maharashtratil Phulpakhre (Butterflies of Maharashtra) (Marathi) (2012, 2016)
eBook
Maharashtratil 100 Samanya Pakshi (Marathi) (2015)
Important Responsibilities
Vice Chairperson: BirdLife International, Asia Division.
Coordinator: Indian Bird Conservation Network (IBCN-BNHS).
National Co-coordinator: Asian Waterbird Census
Executive President: Maharashtra Pakshimitra Sanghatana
Ph.D. Supervisor, Zoology, Mumbai University, Mumbai
Editor, “Mistnet” Quarterly Newsletter of IBCN-BNHS.
Member, Editorial Board: Newsletter for Birdwatchers, Bangalore