Mit12 006jf22 Lec2-3
Mit12 006jf22 Lec2-3
D. H. Rothman, MIT
September 15, 2022
Contents
1 Flows in one dimension 1
1.1 Fixed points and stability . . . . . . . . . . . . . . . . . . . . 2
1.1.1 General statements . . . . . . . . . . . . . . . . . . . . 2
1.1.2 Example: logistic growth . . . . . . . . . . . . . . . . . 3
1.2 Linear stability analysis . . . . . . . . . . . . . . . . . . . . . 5
The simplest dynamical systems concern the evolution of only one variable.
Here we consider flows in which the the dynamics
ẋ = f (x)
1
1.1 Fixed points and stability
f (x) = x2 − 1.
O X
The sign of f (x) determines the directions of the flow of x; we signify this by
arrows.
f (x) intersects the x-axis when f (x) = 0. These are called fixed points (or
equilibrium solutions) of the system. We shall denote these by x⇤ . Here we
have
x⇤ = ±1.
2
1.1.2 Example: logistic growth
Moreover it can even become negative, meaning that the death rate exceeds
the birth rate.
i K N
3
The point at which r(N ) = 0 is a special population size which neither grows
nor decays. It corresponds to N = K, where
K = carrying capacity.
In most cases we can’t really know the shape of r(N ), but the notion of a
carrying capacity remains reasonable.
I N
Then ◆ ✓
N
r(N ) = r 1 −
K
and our growth model now reads
✓ ◆
dN N
= rN 1 − , (1)
dt K
known as the logistic equation.⇤
The logistic equation can be solved exactly but in this course we are generally
interested in qualitative analyses that lead to physical (and mathematical)
insight.
Here, qualitative analysis begins by plotting dN/dt vs. N , i.e., we plot the
RHS of the logistic equation:
N
0
in to
We consider only N ≥ 0 since populations cannot be negative.
⇤
The Belgian mathematician Verhulst published and named it in 1838, but he gave no reason for the
name.
4
The fixed points occur where Ṅ = 0. We find
N1⇤ = 0, N2⇤ = K.
The slope of the curve or the directions of flow tell us that N1⇤ = 0 is unstable
N2⇤ = K is stable.
Indeed, we see that as long as we initiate growth with N (0) > 0 , the popu-
lation always evolves to the carrying capacity.
É I e
5
Now note that
d
⌘˙ = (x − x⇤ )
dt
= ẋ
= f (x)
= f (x⇤ + ⌘).
Expand f (x⇤ + ⌘):
f (x⇤ + ⌘) = f (x⇤ ) + ⌘f 0 (x⇤ ) + O(⌘ 2 ).
Since x⇤ is a fixed point, f (x⇤ ) = 0. Then
f (x⇤ + ⌘) = ⌘f 0 (x⇤ ) + O(⌘ 2 ).
Assuming that f 0 (x⇤ ) 6= 0, the terms of O(⌘ 2 ) are negligible and we have
⌘˙ = ⌘f 0 (x⇤ )
This simple linear equation shows us what we determined earlier from the
slope of f (x) in our graphs:
But we now learn something else: the characteristic timescale of the expo-
nential growth or decay is 1/|f 0 (x⇤ )|.
7
x
Hoax had x x
Nco no r 0
The bifurcation occurs at r = 0.
8
X X X
xno
no mo
Max r 0
ẋ = r − x − e−x
The bifurcation occurs where the graphs of r−x and e−x intersect tangentially.
This requires that
r − x = e−x
and that their slopes are equal:
d d −x
(r − x) = e .
dx dx
The second condition gives
−1 = −e−x ) x=0
We now show why the quadratic form is general (and is therefore called a
9
normal form). Near the bifurcation, we write
e−x
ẋ = r − x − ✓ ◆
x2
= r−x− 1−x+ + ...
2
x2
= r−1− + ...
2
This has the same form as ẋ = r − x2 , and could agree exactly by rescaling
r and x.
10
2.1.2 Example: oxygenation of the atmosphere
ẋ = inputs − outputs
Today’s level of O2 is stable. Conceptually that means there are sources and
sinks that balance in a stable way [2]:
sink
rate
Edt
X
son e
La
There was essentially no O2 in the atmosphere until about 2.5 billion years
ago, when oxygen-producing photosynthesis developed.
How, then, could it have increased if the stability argument just given is
correct?
0
2
source
sink
0
0
˜ ° [O2] ˜
ox
Then, as the sink function weakened, two new fixed points would appear (via
a first saddle-node bifurcation):
11
source − sink
d[O ] / dt ˜
rate °
0
2
source
sink
0
0
˜ ° [O2] ˜
ox
Further weakening of the sink then causes the low-O2 fixed point to collide
with the unstable fixed point via a second saddle-node bifurcation, leaving
only the high-O2 state:
source − sink
d[O2] / dt ˜
rate °
source
sink
0
0
˜ox ° [O2] ˜
For further details and evidence that such a picture may be correct, see
Ref. [3].
In some problems a fixed point must always exist. For example, in the logistic
equation, the zero-population fixed point must always be present.
However the stability of the permanent fixed point may change as a parameter
is varied.
The normal form of the transcritical bifurcation accounts for this possibility.
Its normal form reads
ẋ = rx − x2
This looks like the logistic equation, but now x and r are no longer constrained
to be positive.
Graphically, we have
12
Strogatz [1], Fig. 3.2.1 See image credit on Page 19.
⇤
We see that the fixed point x = 0 always occurs, but as r increases past zero
the unstable fixed point x⇤ = r for r < 0 becomes stable when r > 0.
Many physical problems exhibit symmetry, such that our one dimensional
dynamics are invariant under x ! −x. Examples:
Such symmetry leads to a pitchfork bifurcation. There are two types: super-
critical and subcritical.
13
2.3.1 Supercritical pitchfork bifurcation
When r = 0, the origin is still stable, but only weakly because the linear term
has vanished. Thus there is no longer an exponential decay towards x⇤ = 0.
This is called critical slowing down.
When r > 0, the origin becomes unstable and two new fixed points occur, at
p
x⇤ = ± r. Both are stable.
14
potential V (x). We define the potential by
dV
ẋ = −
dx
so that x tends to go in the direction of smaller V (i.e., downhill), without
inertia, at a rate proportional to the slope of the potential.
We can also now better appreciate the notion of critical slowing down: at
r = 0 the potential is nearly flat near the origin, so fluctuations relax very
slowly.
In the supercritical case, the cubic term is stabilizing: its sign is opposite
that of the instability caused by the linear term.
To see what this means, consider x very small; then the nonlinear term is
negligible so that, for r > 0,
ẋ ⇠ rx ) x ⇠ ert .
15
But eventually the cubic term becomes significant. If we write the supercrit-
ical normal form as ✓ ◆
x2
ẋ = rx 1 − ,
r
p
we see that “saturation” of the instability occurs when x = ± r, i.e., at a
fixed point.
However, nothing prevents the lowest-order nonlinear term from also being
destabilizing. Thus we could have
ẋ = rx + x3 .
The flows are now flipped compared to previously:
x x x
from
to
x
MCO no no
⇤
p
The non-zero fixed points x = ± −r are now unstable, and they exist only
for r < 0. (Thus we see some reason for the name “subcritical.”)
16
Although things now become somewhat complicated algebraically, the e↵ect
of the damping term must create a new set of stable fixed points above the
dashed lines.
0.10
0.1
0.05
-0.05
-0.1
-0.10
-0.2
r=0 r=0.1
x x
0.2
0.2
0.1
0.1
-0.1
-0.1
-0.2
-0.2
• When rs < r < 0, both the origin and the new large-amplitude fixed
points are stable. The initial condition determines the final outcome.
Thus the origin is stable to small perturbations, but not to large pertur-
bations.
• Jumps. When r is increased to r = 0, the system jumps to a large-
amplitude fixed point.
17
• Hysteresis. If r is then decreased, the system stays on the large ampli-
tude branch until it reaches rs , and then it jumps back to the origin.
The dynamics are not reversible.
x
0
rs 0 r
Because the jump can be quite large, subcritical bifurcations are sometimes
called hard or dangerous, while supercritical bifurcations are called soft or
safe.
And in the analogy with phase transitions, supercritical bifurcations are re-
lated to continuous or second-order phase transitions (e.g., the ferromagnetic
transition), while subcritical bifurcations are discontinuous or first-order (e.g.,
the freezing of water into ice).
References
1. Strogatz, S. Nonlinear Dynamics and Chaos: With Applications to Physics,
Biology, Chemistry, and Engineering (CRC Press, 2018).
18
2. Holland, H. D. The Chemistry of the Atmosphere and Oceans (John Wiley
& Sons, New York, 1978).
3. Shang, H., Rothman, D. H. & Fournier, G. P. Oxidative metabolisms
catalyzed Earth’s oxygenation. Nature communications 13, 1–9 (2022).
Image Credit
Images on Pages 8–10 and 13–18 from Strogatz, S. Nonlinear Dynamics and Chaos: With
Applications to Physics, Biology, Chemistry, and Engineering (CRC Press, 2018) © Informa
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19
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