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The document provides links to download various editions of textbooks related to plant biochemistry, including the fourth edition of 'Plant Biochemistry' by Hans-Walter Heldt and Birgit Piechulla. It outlines the focus of the textbook on plant metabolism, biotechnology, and the importance of plants in agriculture and industry. The content emphasizes a balance between general and specialized biochemistry, aiming to facilitate understanding of plant functions and applications.

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Plant Biochemistry 4ed Edition Heldt H.-W. Digital
Instant Download
Author(s): Heldt H.-W., Piechulla B.
ISBN(s): 9780123849861, 0123849861
Edition: 4ed
File Details: PDF, 4.97 MB
Year: 2010
Language: english
Plant Biochemistry

Fourth edition
Plant
Biochemistry
Hans-Walter Heldt
Birgit Piechulla
in cooperation with Fiona Heldt

Translation of the 4th German edition

AMSTERDAM • BOSTON • HEIDELBERG • LONDON


NEW YORK • OXFORD • PARIS • SAN DIEGO
SAN FRANCISCO • SINGAPORE • SYDNEY • TOKYO
Academic Press is an imprint of Elsevier
Academic Press is an imprint of Elsevier
32 Jamestown Road, London NW1 7BY, UK
30 Corporate Drive, Suite 400, Burlington, MA 01803, USA
525 B Street, Suite 1800, San Diego, CA 92101-4495, USA

Fourth edition 2011

Translation © Elsevier Inc.


Translation from the German language edition:
Pflanzenbiochemie by Hans-Walter Heldt and Birgit Piechulla
Copyright © Spektrum Akademischer Verlag Heidelberg 2008
Spektrum Akademischer Verlag is an imprint of Springer-Verlag GmbH
Springer-Verlag GmbH is a part of Springer ScienceBusiness Media
All Rights Reserved

No part of this publication may be reproduced, stored in a retrieval system


or transmitted in any form or by any means electronic, mechanical, photocopying,
recording or otherwise without the prior written permission of the publisher
Permissions may be sought directly from Elsevier’s Science & Technology Rights
Department in Oxford, UK: phone (44) (0) 1865 843830; fax (44) (0) 1865 853333;
email: [email protected]. Alternatively, visit the Science and Technology Books
website at www.elsevierdirect.com/rights for further information

Notice
No responsibility is assumed by the publisher for any injury and/or damage to persons
or property as a matter of products liability, negligence or otherwise, or from any use or
operation of any methods, products, instructions or ideas contained in the material herein.
Because of rapid advances in the medical sciences, in particular, independent verification of
diagnoses and drug dosages should be made

British Library Cataloguing-in-Publication Data


A catalogue record for this book is available from the British Library

Library of Congress Cataloging-in-Publication Data


A catalog record for this book is available from the Library of Congress

ISBN : 978-0-12-384986-1

For information on all Academic Press publications


visit our website at elsevierdirect.com

Typeset by MPS Limited, a Macmillan Company, Chennai, India


www.macmillansolutions.com

Printed and bound in United States of America

10 11 12 13 14 15 10 9 8 7 6 5 4 3 2 1
Dedicated to my teacher, Martin Klingenberg
Hans–Walter Heldt
Preface

The present textbook is written for students and is the product of more than
three decades of teaching experience. It intends to give a broad but concise
overview of the various aspects of plant biochemistry including molecular
biology. We attached importance to an easily understood description of the
principles of metabolism but also restricted the content in such a way that a
student is not distracted by unnecessary details. In view of the importance
of plant biotechnology, industrial applications of plant biochemistry have
been pointed out, wherever it was appropriate. Thus special attention was
given to the generation and utilization of transgenic plants.
Since there are many excellent textbooks on general biochemistry, we
have deliberately omitted dealing with elements such as the structure and
function of amino acids, carbohydrates and nucleotides, the function of
nucleic acids as carriers of genetic information and the structure and func-
tion of proteins and the basis of enzyme catalysis. We have dealt with top-
ics of general biochemistry only when it seemed necessary for enhancing
understanding of the problem in hand. Thus, this book is in the end a com-
promise between a general and a specialized textbook.
To ensure the continuity of the textbook in the future, Birgit Piechulla
is the second author of this edition. We have both gone over all the chap-
ters in the fourth edition, HWH concentrating especially on Chapters
1–15 and BP on the Chapters 16–22. All the chapters of the book have been
thoroughly revised and incorporate the latest scientific knowledge. Here
are just a few examples: the descriptions of the metabolite transport and
the ATP synthase were revised and starch metabolism and glycolysis were
dealt with intensively. The descriptions of the sulfate assimilation and vari-
ous aspects of secondary assimilation, especially the isoprenoid synthesis,
have been expanded. Because of the rapid advance in the field of phytohor-
mones and light sensors it was necessary to expand and bring this chapter
up to date. The chapter on gene technology takes into account the great
advance in this field. The literature references for the various chapters have
been brought up to date. They relate mostly to reviews accessible via data
banks, for example PubMed, and should enable the reader to attain more
detailed information about the often rather compact explanations in the

xxi
xxii Preface

textbook. In future years these references should facilitate opening links to


the latest literature in data banks.
I (HWH) would like to express my thanks to Prof. Ivo Feussner, direc-
tor of the biochemistry division – as emeritus, I had the infrastructure of
the division at my disposal, an important precondition for producing this
edition.
Our special thanks go to the Spektrum team, particularly to Mrs. Merlet
Behncke-Braunbeck who encouraged us to work on this new edition and gave
us many valuable suggestions. We also thank Fiona Heldt for her assistance.
We are very grateful to the Elsevier team for their friendly and very
fruitful cooperation. Our thanks go in particular to Kristi Gomez for the
vast effort she invested in advancing the publication of our translation. We
also thank Pat Gonzalez and Caroline Johnson for their thoughtful sup-
port for our ideas about the layout of this book and their excellent work on
its production.
Once again many colleagues have given us valuable suggestions for the
latest edition. Our special thanks go to the colleagues listed below for criti-
cal reading of parts of the text and for information, material and figures.

Prof. Erwin Grill, Weihenstephan-München


Prof. Bernhard Grimm, Berlin
Steven Huber, Illinois, USA
Wolfgang Junge, Osnabrück
Prof. Klaus Lendzian, Weihenstephan-München
Prof. Gertrud Lohaus, Wuppertal
Prof. Katharina Pawlowski, Stockholm
Prof. Sigrun Reumann, Stavanger
Prof. David. G. Robinson, Heidelberg
Prof. Matthias Rögner, Bochum
Prof. Norbert Sauer, Erlangen
Prof. Renate Scheibe, Osnabrück
Prof. Martin Steup, Potsdam
Dr. Olga Voitsekhovskaja, St. Petersburg

We have tried to eradicate as many mistakes as possible but probably


not with complete success. We are therefore grateful for any suggestions
and comments.

Hans-Walter Heldt
Birgit Piechulla
Göttingen and Rostock, May 2008 (German edition)
July 2010 (Translation)
Introduction

Plant biochemistry examines the molecular mechanisms of plant life. One


of the main topics is photosynthesis, which in higher plants takes place
mainly in the leaves. Photosynthesis utilizes the energy of the sun to syn-
thesize carbohydrates and amino acids from water, carbon dioxide, nitrate
and sulfate. Via the vascular system a major part of these products is trans-
ported from the leaves through the stem into other regions of the plant,
where they are required, for example, to build up the roots and supply them
with energy. Hence the leaves have been given the name “source,” and the
roots the name “sink.” The reservoirs in seeds are also an important group
of the sink tissues, and, depending on the species, act as a store for many
agricultural products such as carbohydrates, proteins and fat.
In contrast to animals, plants have a very large surface, often with very
thin leaves in order to keep the diffusion pathway for CO2 as short as pos-
sible and to catch as much light as possible. In the finely branched root
hairs the plant has an efficient system for extracting water and inorganic
nutrients from the soil. This large surface, however, exposes plants to all
the changes in their environment. They must be able to withstand extreme
conditions such as drought, heat, cold or even frost as well as an excess
of radiated light energy. Day to day the leaves have to contend with the
change between photosynthetic metabolism during the day and oxida-
tive metabolism during the night. Plants encounter these extreme changes
in external conditions with an astonishingly flexible metabolism, in which
a variety of regulatory processes take part. Since plants cannot run away
from their enemies, they have developed a whole arsenal of defense sub-
stances to protect themselves from being eaten.
Plant agricultural production is the basis for human nutrition. Plant
gene technology, which can be regarded as a section of plant biochemis-
try, makes a contribution to combat the impending global food shortage
due to the enormous growth of the world population. The use of envi-
ronmentally compatible herbicides and protection against viral or fungal
infestation by means of gene technology is of great economic importance.
Plant biochemistry is also instrumental in breeding productive varieties of
crop plants.

xxiii
xxiv Introduction

Plants are the source of important industrial raw material such as fat
and starch but they are also the basis for the production of pharmaceutics.
It is to be expected that in future gene technology will lead to the extensive
use of plants as a means of producing sustainable raw material for indus-
trial purposes.
The aim of this short list is to show that plant biochemistry is not only
an important field of basic science explaining the molecular function of a
plant, but is also an applied science which, now at a revolutionary phase of
its development, is in a position to contribute to the solution of important
economic problems.
To reach this goal it is necessary that sectors of plant biochemistry such
as bioenergetics, the biochemistry of intermediary metabolism and the sec-
ondary plant compounds, as well as molecular biology and other sections
of plant sciences such as plant physiology and the cell biology of plants,
co-operate closely with one another. Only the integration of the results and
methods of working with the different sectors of plant sciences can help
us to understand how a plant functions and to put this knowledge to eco-
nomic use. This book will try to describe how this could be achieved.
Since there are already very many good general textbooks on biochem-
istry, the elements of general biochemistry will not be dealt with here and it
is presumed that the reader will obtain the knowledge of general biochem-
istry from other textbooks.
1
A leaf cell consists of several
metabolic compartments

In higher plants photosynthesis occurs mainly in the mesophyll, the


chloroplast-rich tissue of leaves. Figure 1.1 shows an electron micrograph
of a mesophyll cell and Figure 1.2 shows a schematic presentation of the
cell structure. The cellular contents are surrounded by a plasma membrane

Figure 1.1 Electron


micrograph of mesophyll
tissue from tobacco. In
most cells the large central
vacuole is to be seen (v).
Between the cells are the
intercellular gas spaces
(ig), which are somewhat
enlarged by the fixation
process. c: chloroplast; cw:
cell wall; n: nucleus; m:
mitochondrion. (By D. G.
Robinson, Heidelberg.)

1
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