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THE TRACKER’S
FIELD GUIDE
A Comprehensive Manual for
Animal Tracking

Third Edition

JAMES C. LOWERY
An imprint of Globe Pequot, the trade division of
The Rowman & Littlefield Publishing Group, Inc.
4501 Forbes Blvd., Ste. 200
Lanham, MD 20706
www​.rowman​.com
Falcon and FalconGuides are registered trademarks and Make Adventure Your Story is a
trademark of The Rowman & Littlefield Publishing Group, Inc.

Distributed by NATIONAL BOOK NETWORK

Copyright © 2022 James C. Lowery

All photos by James C. Lowery unless otherwise noted.

Maps by Melissa Baker © The Rowman & Littlefield Publishing Group, Inc

All rights reserved. No part of this book may be reproduced in any form or by any
electronic or mechanical means, including information storage and retrieval systems,
without written permission from the publisher, except by a reviewer who may quote
passages in a review.

British Library Cataloguing in Publication Information available

Library of Congress Cataloging-­in-­Publication Data available

ISBN 978-1-4930-6703-9 (paper: alk paper)

ISBN 978-1-4930-6704-6 (electronic)

The paper used in this publication meets the minimum requirements of American
National Standard for Information Sciences—Permanence of Paper for Printed Library
Materials, ANSI/NISO Z39.48-1992.

The authors and The Rowman & Littlefield Publishing Group, Inc. assume no liability
for accidents happening to, or injuries sustained by, readers who engage in the
activities described in this book.
To those who devote their lives to habitat protection
CONTENTS

Acknowledgments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . vii
1: Into the Animal’s World . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1
2: Clear Print Identification . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
3: Track Pattern Basics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9
4: Soil Movement and Track Identification . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
5: Track Pattern Interpretation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
6: Using Pressure Releases for Interpretation . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26
7: Mammal Pages . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 30

CATS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 33
Bobcat . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 34
Domestic Cat . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 42
Mountain Lion . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 46

DOGS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55
Coyote . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 56
Domestic Dog . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 64
Gray Fox . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68
Gray Wolf . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 76
Kit Fox . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 82
Red Fox . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 90

RABBITS AND HARES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 99

Black-­Tailed Jackrabbit . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 100
Cottontail . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 108
Snowshoe Hare . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 116
White-­Tailed Jackrabbit . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 122

RODENTS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 127
Antelope Squirrel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 128
Beaver . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 132
Chipmunk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 140
Fox Squirrel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 148
Gray Squirrel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 154
 Contents

Ground Squirrel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 162

Kangaroo Rat . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 170
Marmot/Woodchuck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 178
Mice . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 184
Peromyscus Mice (Peromyscus Genus) . . . . . . . . . . . . . . . . . . . . . . . . . . 187
Harvest Mice (Reithrodontomys Genus) . . . . . . . . . . . . . . . . . . . . . . . . . 188
Grasshopper Mice (Onychomys Genus) . . . . . . . . . . . . . . . . . . . . . . . . . 189
Pocket Mice (Chaetodipus and Perognathus Genus) . . . . . . . . . . . . . 190
Muskrat . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 192
Pocket Gopher . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 198
Porcupine . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 202
Red Squirrel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 210
Vole . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 216
Woodrat . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 220

WEASELS AND SKUNKS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 229

Badger . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 230
Fisher . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 238
Marten . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 242
Mink . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 250
River Otter . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 256
Spotted Skunk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 262
Striped Skunk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 270
Weasel . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 278

RACCOONS, OPOSSUMS, AND BEARS . . . . . . . . . . . . . . . . . . . . . . . 287

Raccoon . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 288
Ringtail . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 296
Opossum . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 304
Black Bear . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 310
Grizzly Bear . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 318

INSECTIVORE . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 323
Shrew . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 324

UNGULATES . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 327
Bighorn Sheep . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 328
Burro . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 336

v
Contents

Domestic Cow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 338

Elk . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 340
Feral Pig . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 348
Llama . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 350
Moose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 352
Mule Deer/Black-­Tailed Deer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 356
Pronghorn . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 364
White-­Tailed Deer . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 370
Appendix A: Mammal List . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 376
Appendix B: Recommended Reading . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 380
Appendix C: Visualizing Home-­Range Sizes . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 396
Appendix D: Resources . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 398
Glossary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 399
Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 402
Clear Print Summary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 404
Track Pattern Summary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 406
About the Author . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 408

vi
ACKNOWLEDGMENTS

Threads from every part of this book lead back to Tom Brown Jr., who intro-
duced me to tracking and who showed that the hard-­to-­imagine is possible.
The complete dedication to the learning process and the honoring of the
track’s maker above all are lessons I will always carry with me.

For the third edition’s conversion and improvement of photos, I am indebted

to help and coaching from Rhanna Nyman; she and Rowan Nyman are also
responsible for cover design elements. I am very thankful for Kim Cabrera’s
help with the Resources appendix stemming from her vital role connecting
trackers with one another.

Doug Gaulke made the book’s completion possible through his substantial
help with illustrations and knowledge about mammals outside my area, such
as the gray wolf and white-­tailed deer. Paul Rezendes, Mark Elbroch, Matt
Gray, Renee Robison, and Dick Newell also contributed essential photos.
Marge Erickson merged her extensive tracking background with professional
editing skills in proofreading the manuscript.

Thanks go to the first edition manuscript reviewers for their suggestions

and encouragement: Don Mitchell, Wanda DeWaard, Roseann and Jonathan
Hanson, Carl Olmstead, and Steve Montgomery. Tom Lowery designed an
attractive and flexible layout concept for the original book development.

Invaluable help in field research as I sought tracks of specific mammals was

provided by Curtis Bjurlin and Alexander Brown of the Endangered Species
Recovery Program at California State University, Stanislaus, who released
grasshopper mice and pocket mice onto my tracking boards; Patty Quickert
and Laura Patterson of the California Department of Water Resources, who
did the same with harvest mice; and Elizabeth Jozwiak of the Kenai National
Wildlife Refuge, who helped with snowshoe-­hare tracks and signs. Alice Koch
helped with pronghorn, badger, and kit fox tracks on the Carrizo Plain and
Larry Lambrecht helped find elk tracks in Arizona.

I am indebted to the owners and managers of the natural areas where we

have conducted classes and collected track information for this book. The
importance of Windy Springs Preserve, and my appreciation of its late owner
and steward Tom Drummond, cannot be overstated. Also I acknowledge the

vii
Acknowledgments

Desert Studies Center, Zzyzx, California; the University of California James

Reserve; the Dunes Center in Guadalupe, California; Wind Wolves Preserve;
the Carrizo Plain National Monument; the Palos Verdes Peninsula Land Con-
servancy; and Lone Juniper Ranch in Gorman, California. Special thanks go
to Steve Martin and the staff of Working Wildlife, where we were able to con-
duct animal-­movement workshops and study tracks of captive wild animals,
and to Rose Berger of SunWolf Farms for her very professional and enthusi-
astic help with horses and their gaits.

The groundwork laid by dedicated tracking teachers over the past thirty years
has benefited all of us who study and teach the art. I would like to acknowl-
edge especially the pioneering work of Tom Hanratty, Charles Worsham, Jim
Halfpenny, Paul Rezendes, Jon Young, Susan Morse, and Mark Elbroch. More
recently the contributions of CyberTracker evaluators and instructors toward
expanding the tracking audience and its expertise continues to be especially
important.

I cannot list all of the dedicated Earth Skills tracking students who accom-
panied us as we learned. Many of these people field-­tested early drafts of this
guide in classes and gave suggestions and encouragement. You know who
you are—thank you. Our First Nation friends Robin, Terry, and Barbara,
along with their extended family, helped to keep us humble and thankful.
Scott Cunningham’s music was more important than he knows.

I am grateful to Globe Pequot Press and Falcon Guides for continuing to

make this book widely available.

Finally, the sacrifice and support I received from my wife, Mary, toward this
book cannot possibly be described and can be understood only by those who
know her. From carefree exploration to times of crushing focus, she was my
constant companion. She is a true tracker because she always finds what is
necessary. This project and my life have been blessed by her.

viii
1
INTO THE ANIMAL’S WORLD

If you are already a tracker, you know what novice trackers will learn in their
first day: The animal whose tracks you just identified is going to call you out.
Where am I going, it will say. Why am I traveling alone? Why did I bypass this
perfectly good food? How recently was I here? Why did I take this side trail?

The tracks won’t answer everything, but they do tell you more than you
probably think. Mood, posture, speed, and personality. The reason for a
choice of a bedding site. The current place in this year’s rearing of young.
Deliberate avoidance of another of its species. An exploratory urge or hungry
focus.

Most importantly, the tracks begin the conversation between you and the
animal, the animal leading the way because it’s the one with the answers.
You don’t need to be an accomplished tracker to hold up your end of the
conversation, but it’s best to know how to read what else is in the tracks
besides the identity of its maker. And, it’s essential to bring in biology and
behavior too.

With these principles in mind—learned from my own fieldwork as well as

from teaching hundreds of tracking workshops—I wrote this book to help
you identify tracks confidently and efficiently, and then to step into the
animal’s world toward a rich and often surprising journey of discovery.

There are a lot of tools here: extensive notes about the biology of each animal,
many examples of track interpretation, and “Track Windows” suggesting
how your exploration can access an animal’s unique personality and biology.
If you are a beginning tracker, you will grow into some of these tools over
time. If you are an advanced tracker, you will find useful ones not found in
other references.

Remember that the opportunity to interact with the animal you are tracking,
whether it be a chipmunk or an elk, is special and sacred. The animal passed
here, in this way, only once. It left behind the signs of its body language, signs
of choices about food and travel, evidence of when it rested and where it ran.
Each of these threads draws inward to the animal’s mood and outward to its

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