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THE FACTS ON FILE
MARINE
SCIENCE
HANDBOOK
SCOTT MCCUTCHEON
and
BOBBI MCCUTCHEON
The Facts On File Marine Science Handbook
Copyright © 2003 by Scott McCutcheon and Bobbi McCutcheon
All rights reserved. No part of this book may be reproduced or utilized in any
form or by any means, electronic or mechanical, including photocopying,
recording, or by any information storage or retrieval systems, without
permission in writing from the publisher. For information contact:
McCutcheon, Scott.
The Facts on File marine science handbook / Scott McCutcheon and Bobbi
McCutcheon.
p. cm. — (the Facts on File science handbooks)
Includes bibliographical references and index.
ISBN 0-8160-4812-6 (acid-free paper)
1. Marine sciences — Handbooks, manuals, etc. I. McCutcheon, Bobbi.
II. Title, III. Series.
GC24.M43 2003
551.46 — dc21 2003000275
Facts On File books are available at special discounts when purchased in bulk
quantities for businesses, associations, institutions, or sales promotions.
Please call our Special Sales Department in New York at 212/967-8800 or
800/322-8755.
VB Hermitage 10 9 8 7 6 5 4 3 2 1
Acknowledgments iv
Preface iv
Introduction vi
APPENDIXES:
Prefixes and Suffixes 258
Recommended Reading
and Useful Websites 260
INDEX 263
ACKNOWLEDGMENTS
We the authors would like to thank the National Oceanic and Atmospheric
Administration (NOAA) and their staff members for generously providing
much of the information and many of the photos herein.
PREFACE
Who are we? Where do we come from? These are two of humankind’s
most asked questions. Though scientists differ on the answer, one that
still yearns to be solved irrefutably, there is one fact we know with
certainty: where there is water there is life. Marine science is the
ultimate requisite when exploring and understanding the world in which
we live. Within our solar system, Earth is the only body to have not only
enough water to support life, but also enough to foster virtually millions
of species, many of which are still waiting to be revealed.
One finds it fascinating that we as human beings live each day above the
unassuming seas while beneath them in reality lies a hidden ecosystem
teeming with creatures and vegetation so bizarre they really seem to be
inhabitants of another world. Some consider space the final frontier; yet
the oceans of Earth are just as mysterious, and as far as we have come in
understanding them, there is still so much to be discovered.
iv
Preface
or the moons of Jupiter, we were to find liquid water, and with it perhaps
ancient or newly forming life, it would undoubtedly generate a staffed
mission. What if the most important person on that future mission were
the marine biologist? What if that biologist were you? But how can we
begin to understand water-generated life on other worlds without first
grasping the secrets of our own?
This primer on marine science is the perfect start. Within these pages
designed especially for you can be found a selected list of easy-to-follow
terms, descriptions, drawings, biographies of the great people who have
made historic contributions in marine science, a chronology of
exploration and research, and straightforward charts to augment the
terms and definitions.
Preface
v
INTRODUCTION
GLOSSARY
This list contains almost 1,500 entries: alphabetically listed terms and
their definitions used in the field of marine science, often accompanied
by a graph or chart to clarify meanings. A well-equipped and easily
understood glossary within arm’s reach is essential for any field of study.
BIOGRAPHIES
Throughout the history of the systematic study of the oceans, many great
people have stood on the shoulders of their eminent predecessors to
broaden knowledge about our marine environment. These early to
modern sailors and scientists began the process that allows today’s mind
to understand more fully our greatest nurturer of life, the oceans. Listed
here are close to 250 of the most renowned of those people, along with
many less familiar individuals who have earned recognition for their
selfless efforts and principal contributions in the study of marine
science.
CHRONOLOGY
On an evolutionary scale, the discovery of science and technology has
evolved on Earth in the twinkle of an eye, but in human terms, for the
most part, it has not been what anyone could call fast paced. Patience
must sometimes be employed; yet the rewards are great, and each
exciting breakthrough inevitably causes change, however gradual.
Nearly 200 entries are listed here, providing a general historical account
of oceanic explorations and marine research spanning 6,000 years,
events that have helped revolutionize civilized humanity’s understanding
of the sea.
vi
SECTION ONE
GLOSSARY
1
GLOSSARY abaft – acidic
abaft Behind. A nautical term meaning toward the stern (rear) of a boat.
abalone A primitive, univalve gastropod found on the rocky shores of warm
and temperate seas, except the western Atlantic Ocean. We value
them for their large, edible foot and the mother-of-pearl lining in
their shells, which commonly is used to make decorative items such
as jewelry. Abalones constitute the family Haliotidae, in the order
Archaeogastropoda, class Gastropoda, phylum Mollusca, kingdom
Animalia.
abdomen In a TETRAPOD, the portion of its body between the thorax and
pelvic girdle. In an ARTHROPOD, the region of its body behind the
thorax. It contains the visceral (internal) organs.
abeam A nautical term meaning at a right angle to a ship’s keel, yet not on
the ship.
abiotic The nonexistence of life, or nonbiological.
abyssal hills Low, undersea hills characteristic only of the deep sea. Mainly,
they rise from about 10 to 750 feet (3–228 m) high and range
anywhere from a few hundred feet to a few miles wide.
abyssal plain Also known as basins or ocean basins, these are flat expanses of
the deep seafloor that slope less than one part in 1,000 and typically
exist between the base of abyssal hills and the continental slope.
Many basins and plains are identified in this book.
abyssal plain
abyssopelagic zone The lightless area of open ocean beneath the
BATHYPELAGIC ZONE, existing at about 12,000 to 20,000 feet
(3,657–6,096 m). Examples of sea creatures adapted to such depths
are deep-water eels, such as gulpers; deep-sea anglers; and
stomiatoids.
acanthodians A group of the earliest known true jawed fishes from the Lower
Silurian to the Lower Permian period. They were sharklike and had
spines on their bodies.
acetosporan A simple PROTOZOAN, parasitic on marine invertebrates, that is
characterized as having multicellular spores. It can cause diseases
and mass mortality.
acicula Needlelike supporting bristle existing in the PARAPODIA of some
POLYCHAETES (worms). It can also exist in the form of crystals of
abyssopelagic zone certain minerals.
acidic A term descriptive of a substance having acidlike qualities and
possessing a relatively large number of hydrogen ions; having a pH
less than 7.0.
2
acoustic oceanography – aerobic GLOSSARY
3
GLOSSARY aeroembolism – Alboran Sea
Alaska, Gulf of A wide body of water centered at about 58° N and 145° W. It
is bordered by Alaska on all sides but the south, which is open to the
North Pacific Ocean and therefore subject to all consequences of the
weather. Despite its being battered by continual storms and
occasional tsunamis, it is a region heavily fished.
Alboran Sea Part of the western MEDITERRANEAN SEA centered at about 4° W
and 36° N. Morocco borders it to the south, Spain to the north, and it
4
Aleutian Current – alluvial GLOSSARY
5
GLOSSARY alluviation – amphipod
6
ampullae of Lorenzini – Angiospermae GLOSSARY
They are in the order Amphipoda below the phylum Arthropoda; the
majority are marine, but a few species are freshwater.
ampullae of Lorenzini Pores forming an electrical receptor system, present in a
number of cartilaginous fishes, that serve to detect weak electrical
fields generated by other living creatures, even when they are hidden.
Amundsen Sea An arm of the Southern Ocean, it is a marginal sea named
after the Norwegian explorer ROALD AMUNDSEN, which lies centered
at about 73° S and 112° W, directly off the coast of Antarctica among
the BELLINGSHAUSEN SEA, to the east; the ROSS SEA, to the west; and
the imaginary Antarctic Circle to the north.
anadromous fish Refers to fish that spend most of their adult lives in the
marine environment but breed in freshwater. SALMON are an example.
Compare to CATADROMOUS and OCEANODROMOUS FISH.
Anadyr Current A surface current that flows along the northwest side of the
BERING SEA and then continues through the BERING STRAIT. This
current is a major factor for depositing biological material into
Chirikov Basin, which lies north of Saint Lawrence Island, and thus
the basin is an active feeding ground for whales.
Anadyr, Gulf of (Anadyrskiy Zaliv) A large embayment of the BERING SEA
centered at about 64° N and 175° W, indenting the extreme eastern
tip of Siberia.
anaerobic Existing in the absence of oxygen.
anal fin In fish, a ventral (belly) fin located behind the anus.
anatomy Pertains to the individual design or the scientific study of the
structure of an entire organism.
Andaman Sea A body of water in the northeast end of the Indian Ocean
centered at about 10° N and 95° E. The Andaman Islands border it to
the west, the Malay Peninsula to the east, Sumatra to the south, and
Burma’s Irrawaddy Delta to the north. It connects to the JAVA SEA by
way of the Malacca Strait.
anemometer An instrument for measuring direction and speed of wind. The
most common anemometer works by a series of cups attached to
horizontal limbs that spin around a vertical support. The rate of
rotation is calibrated to obtain wind speed.
angiosperm A plant whose seeds are produced in an ovary, such as a flower,
fruit, or SEA GRASS, as opposed to a gymnosperm, whose seeds are
not produced in an ovary, such as a conifer.
Angiospermae Phylum of the salt marsh grasses, sea grasses, and mangroves,
within the kingdom Plantae. anemometer
7
GLOSSARY Angola Basin – Antarctic Convergence
Angola Basin A basin of the Atlantic Ocean located west of Africa directly
along the prime meridian at about 15° S longitude. It lies between the
Angola Abyssal Plain, to the south, and the Guinea Ridge, to the
north. Also known as the Buchanan Deep.
Angola Current In the Atlantic Ocean, it is part of a subsurface gyre driven by
the South Equatorial Countercurrent centered at about 13° S and 4° E
at around 1,000 feet (304 m) in depth.
ANGUS (acoustically navigated underwater survey system) This equipment
operated by SCRIPPS INSTITUTION OF OCEANOGRAPHY is essentially
an unstaffed sled with observation cameras. In 1977 it made its first
observation of deep-sea vent communities.
Animalia One of the five taxonomic kingdoms of living organisms whose
members are multicellular, eukaryotic (having a true nucleus), and
heterotrophic. The other kingdoms are Fungi, Monera, Plantae, and
Protozoa. Organisms that ingest food from outside sources are
classified as animals, as opposed to plants, which generate their own
energy through PHOTOSYNTHESIS. Within this section, many marine-
related phyla of the animal kingdom are recognized.
annelid Common name for wormlike invertebrates, such as the marine bristle
worms, that make up the class Polychaeta, under the phylum
Annelida. Recently, scientists discovered a new species of marine
worm living in frozen methane in the deep waters of the Gulf of
Mexico. New discoveries within the marine environment are being
made each year.
Annelida Phylum of the polychaete worms and leeches, within the kingdom
Animalia.
annulus The growth ring in a fish SCALE.
anoxic Deficient in oxygen.
Antarctic Cold climactic regions south of the Antarctic Circle, surrounding the
South Pole. Antarctic means the opposite of Arctic.
Antarctic Circle Latitude line 66.5° south of the equator in Earth’s Southern
Hemisphere. That places it 23.5° north of the South Pole. On June 21
(winter solstice), the Sun never rises above the horizon, and on
December 22 (summer solstice), the Sun never sets below the horizon.
See also ARCTIC CIRCLE.
Antarctic Convergence An area of ocean encircling Antarctica between
latitudes 47° S and 62° S that experiences rapid temperature changes
from current subduction and convergence.
8
Antarctic Current – aquaculture GLOSSARY
Antarctic Current The principal current of the Southern Ocean region, directly
associated with the EAST WIND DRIFT and the WEST WIND DRIFT. The
temperature and salinity effects of this powerful current are global.
Antarctic Ocean The name sometime applied to the SOUTHERN OCEAN, the
fifth and most recently classified division of the WORLD OCEAN.
anterior The head or forward end of an animal.
See also POSTERIOR.
Anthozoa A class of marine COELENTERATES in the phylum Cnidaria
(Coelenterata). The anthozoans are animals with radial symmetry,
such as sea anemones and some corals.
anthracosaur A Paleozoic labyrinthodont amphibian. Labyrinthodonts
typically resembled salamanders or crocodiles. They are considered
to be the earliest TETRAPOD vertebrates.
Antilles Current A clockwise-circulating current of the North Atlantic Ocean.
It is one of the distinct currents of this area’s circulation system,
which includes the NORTH EQUATORIAL CURRENT, GULF STREAM,
North Atlantic Drift, and CANARY CURRENT.
AODC (Australian Oceanographic Data Center) Established within the Royal
Australian Navy in 1964, this center participates in national and
international projects in many branches of marine science.
See also Useful websites in the Appendixes.
AOML (Atlantic Oceanographic and Meteorological Laboratory) Part of the
NOAA (National Oceanic and Atmospheric Administration) network
located in Miami, Florida, whose mission is to conduct research
programs in oceanography, tropical meteorology, atmospheric and
oceanic chemistry, and ACOUSTICS.
See also Useful websites in the Appendixes; NOAA.
aphotic Devoid of light.
aphotic zone Deep, open-ocean region below the DISPHOTIC ZONE, sometimes
called the midnight zone, where no light penetrates and where
PHOTOSYNTHESIS may never occur.
See also EUPHOTIC ZONE.
aplacophora A wormlike MOLLUSK, rarely seen, having small spines that
cover its body. Aplacophora is a Greek word meaning, “no shell.”
apneustic breathing A breathing pattern in marine mammals whereby a number
of rapid breaths alternate with a prolonged pause in breathing.
aquaculture Cultivation of aquatic organisms under controlled conditions.
9
GLOSSARY Aqua-Lung – Arctic Ocean
Arabian Sea A northwest arm of the Indian Ocean centered at around 16° N
and 65° E, bounded by the Arabian Peninsula on the west, the
subcontinent of India on the east, the greater Indian Ocean on the
south, and Iran and Pakistan on the north.
Arafura Sea Centered at about 10° S and 137° E, this is a marginal sea belonging
to the waters around northern Australia and Indonesia. Australia borders
it to the south, Irian Jaya and New Guinea to the north-northeast, the
TIMOR SEA to the west, and the Gulf of Carpentaria to the southeast.
Aral Sea Centered at around 45° N and 60° E, this is a saltwater lake resting
along the border between Uzbekistan and Kazakhstan. It was once part
of the CASPIAN SEA, before the last ice age.
See also TETHYS SEA.
archipelagic apron Common in the Central and South Pacific Ocean, a
broad, fan-shaped or cone-shaped underwater slope(s) forming
against a volcanic seamount(s). In typical development, a volcano
erupts and depresses the surrounding crust; then the depression fills
with lava and new sediment to form the archipelagic apron.
archipelago A group or chain of islands, an example of which is the
Hawaiian Islands or Alaska’s Aleutian Island chain.
Arctic Cold climactic regions north of the ARCTIC CIRCLE, surrounding the
North Pole.
Arctic Circle Latitude line 66.5° north of the equator in Earth’s Northern
Hemisphere. That places it 23.5° south of the North Pole.
See also ANTARCTIC CIRCLE.
Arctic Ocean Ocean area north of North America and Eurasia. It is one of the
five divisions of the WORLD OCEAN and is covered all winter with
pack ice and therefore never totally navigable. Its surface area covers
5,427,000 square miles (14,056,000 km2), one-third of that over
Arctic Ocean continental shelf.
10
Argentine Basin – asexual reproduction GLOSSARY
11
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